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Selden, P.A., Shcherbakov, D.E., Dunlop, J.A. & Eskov, K.Y. (2014). Arachnids from the Carboniferous of Russia and Ukraine, and the Permian of Kazakhstan. Paläontologische Zeitschrift 88: 297–307. doi: 10.1007/s12542-013-0198-9