Larca granulata (Banks, 1891)


Original combination: Garypus granulatus


Taxonomic references

Garypus granulatus

Banks, 1891: 163–164 (as Grypus [sic] granulatus) original description
Banks, 1895a: 9
Coolidge, 1908: 112
Chamberlin, 1921: 191

Larca granulata (Banks):

Chamberlin, 1930: 616, figs 3m-n
Beier, 1932a: 224–225
Roewer, 1937: 268
Hoff, 1945a: 109–110, figs 9–10
Hoff, 1949b: 447–448, figs 26, 27a–d
Hoff and Bolsterli, 1956: 163–164
Hoff, 1958: 15
Weygoldt, 1969a: n/a, fig. 51
Muchmore, 1971a: 90, 92
Nelson, 1975: 282, figs 29–31
Benedict and Malcolm, 1978a: 117
Muchmore, 1981a: 50–51
Zeh, 1987b: 1086
Harvey, 1990d: 157–159, fig. 3
Muchmore, 1990a: 514, figs 18.3, 18.6–18.7
Harvey, 1991a: 242
Snider and Nelson, 1991: 261
Harvey, 1992c: n/a, figs 136–141
Whitford and Steinberger, 2010: 1454 (possible misidentification; not included in distribution)
Benavides, Cosgrove, Harvey and Giribet, 2019: 3, 4, figs 1E, 3, 4

Chernes dentatus (Banks):

Ross, 1944: n/a, fig. 56 (misidentification)

Distribution table

Realm: Nearctic

US (Illinois, Kansas, Michigan, New Hampshire, New York, North Carolina, Pennsylvania, Tennessee, Virginia, West Virginia)
Type locality: Ithaca, Tompkins County, New York, U.S.A.


Locality Lat Lon
1 Ithaca 42.433333 -76.5
Type repository
Zoobank LSID


Banks, N. (1891). Notes on North American Chernetidae. Canadian Entomologist 23: 161–166. -- Show included taxa

Banks, N. (1895a). Notes on the Pseudoscorpionida. Journal of the New York Entomological Society 3: 1–13. -- Show included taxa

Beier, M. (1932a). Pseudoscorpionidea I. Subord. Chthoniinea et Neobisiinea. Tierreich 57: i–xx, 1–258. -- Show included taxa

Benavides, L.R., Cosgrove, J.G., Harvey, M.S. & Giribet, G. (2019). Phylogenomic interrogation resolves the backbone of the Pseudoscorpiones Tree of Life. Molecular Phylogenetics and Evolution 139(106509): 1–14. -- Show included taxa

Benedict, E.M. & Malcolm, D.R. (1978a). Some garypoid false scorpions from western North America (Pseudoscorpionida: Garypidae and Olpiidae). Journal of Arachnology 5: 113–132. -- Show included taxa

Chamberlin, J.C. (1921). Notes on the genus Garypus in North America (Pseudoscorpionida - Cheliferidae). Canadian Entomologist 53: 186–191. -- Show included taxa

Chamberlin, J.C. (1930). A synoptic classification of the false scorpions or chela-spinners, with a report on a cosmopolitan collection of the same. Part II. The Diplosphyronida (Arachnida-Chelonethida). Annals and Magazine of Natural History (10) 5: 1–48, 585–620. -- Show included taxa

Coolidge, K.R. (1908). A list of the North American Pseudoscorpionida. Psyche, Cambridge 15: 108–114. -- Show included taxa

Harvey, M.S. (1990d). Trichobothrial "migration" in diplosphyronid pseudoscorpions. Acta Zoologica Fennica 190: 157–160. -- Show included taxa

Harvey, M.S. (1991a). Catalogue of the Pseudoscorpionida. Manchester University Press, Manchester. -- Show included taxa

Harvey, M.S. (1992c). The phylogeny and classification of the Pseudoscorpionida (Chelicerata: Arachnida). Invertebrate Taxonomy 6: 1373–1435. -- Show included taxa

Hoff, C.C. (1945a). Additional notes on pseudoscorpions from Illinois. Transactions of the Illinois Academy of Science 38: 103–110. -- Show included taxa

Hoff, C.C. (1949b). The pseudoscorpions of Illinois. Bulletin of the Illinois Natural History Survey 24: 407–498. -- Show included taxa

Hoff, C.C. (1958). List of the pseudoscorpions of North America north of Mexico. American Museum Novitates 1875: 1–50. -- Show included taxa

Hoff, C.C. & Bolsterli, J.E. (1956). Pseudoscorpions of the Mississippi River drainage basin area. Transactions of the American Microscopical Society 75: 155–179. -- Show included taxa

Muchmore, W.B. (1971a). Phoresy by North and Central American pseudoscorpions. Proceedings of the Rochester Academy of Science 12: 79–97. -- Show included taxa

Muchmore, W.B. (1981a). Cavernicolous species of Larca, Archeolarca and Pseudogarypus with notes on the genera, (Pseudoscorpionida, Garypidae and Pseudogarypidae). Journal of Arachnology 9: 47–60. -- Show included taxa

Muchmore, W.B. (1990a). Pseudoscorpionida. In: Dindal, D.L. (ed.) Soil biology guide John Wiley and Sons, New York: 503–527. -- Show included taxa

Nelson, S.O., Jr (1975). A systematic study of Michigan Pseudoscorpionida (Arachnida). American Midland Naturalist 93: 257–301. -- Show included taxa

Roewer, C.F. (1937). Chelonethi oder Pseudoskorpione. In: Bronns, H.G. (ed.) Bronn's Klassen und Ordnungen des Tierreichs Akademische Verlagsgesellschaft M.B.H., Leipzig 5(IV)(6)(1): 161–320. -- Show included taxa

Ross, H.H. (1944). How to collect and preserve insects. Illinois Natural History Survey, Divisional Circular 39: 1–55. -- Show included taxa

Snider, R.J. & Nelson, S.O., Jr (1991). Michigan Pseudoscorpionida, a checklist. Michigan Academician 24: 259–263. -- Show included taxa

Weygoldt, P. (1969a). The biology of pseudoscorpions. Harvard University Press, Cambridge, Massachusetts. -- Show included taxa

Whitford, W.G. & Steinberger, Y. (2010). Pack rats (Neotoma spp.): keystone ecological engineers? Journal of Arid Environments 74(11): 1450–1455. -- Show included taxa

Zeh, D.W. (1987b). Aggression, density, and sexual dimorphism in chernetid pseudoscorpions (Arachnida: Pseudoscorpionida). Evolution 41: 1072–1087. -- Show included taxa

Date Type Reference Old value
2022-07-03 New taxonomic reference entry Benavides, Cosgrove, Harvey and Giribet, 2019