Anchigarypus californicus (Banks, 1909)


Original combination: Garypus californicus


Taxonomic references

Garypus californicus

Banks, 1909b: 305 original description
Banks, 1911: 635, fig. 210b
Moore, 1917: 26, fig. 1
Chamberlin, 1921: 190, figs a–d
Moles and Moore, 1921: 8
Chamberlin, 1925a: 330
Chamberlin, 1930: 613, figs 2j, 2o, 2s, 2z, 3p, 3s
Chamberlin, 1931a: n/a, figs 3, 4c, 6c, 8b, 11p, 14c–d, 16n, 17w, 19j, 21d, 24a, 26a, 29f, 40u, 41e–f, 45o, 46l, 47f, 47t, 50f
Beier, 1932a: 219–220, fig. 246
Beier, 1932g: n/a, figs 159, 185
Chamberlin, 1935b: 480
Roewer, 1937: 268
Hoff, 1958: 15
Ricketts and Calvin, 1960: 432
Firstman, 1973: 5, 16, fig. 10
Newell, 1975: 431, plate 105 fig. 2
Roth and Brown, 1976: 128
Schulte, 1976: 119–123, figs 1–2
Lee, 1979a: 4–7, figs 2, 4, 6–8, 30
Lee, 1979b: 13
Poinar, Thomas and Lee, 1985: 400–401, fig. 1
Zeh, 1987b: 1086
Muchmore, 1990a: 514
Harvey, 1991a: 239
Austin, Gibson and Harvey, 1998: 349
Lighton and Joos, 2002a: 345–348, figs 1–5
Lighton and Joos, 2002b: 1–4, figs 1–3
Ceballos, 2004: 428
Benavides, Cosgrove, Harvey and Giribet, 2019: 3, 4, figs 3, 4

Anchigarypus californicus (Banks, 1909):

Harvey, Hillyer, Carvajal and Huey, 2020: 37, 46, figs 1, 2

Distribution table
US (California)
Type locality: Palo Alto, Santa Clara County, California, U.S.A. San Nicolas Island, Ventura County, California, U.S.A.


Locality Lat Lon
1 Palo Alto 37.466666 -122.116666
2 San Nicolas Island 33.25 -119.5
Type repository
  • undefined Syntypes: 2 specimens.
Zoobank LSID


Austin, A.D., Gibson, G.A.P. & Harvey, M.S. (1998). Synopsis of Australian Calymmochilus Masi (Hymenoptera: Eupelmidae), description of a new Western Australian species associated with a pseudoscorpion, and review of pseudoscorpion parasites. Journal of Natural History 32: 329–350. -- Show included taxa

Banks, N. (1909b). Arachnida from Costa Rica. Proceedings of the Academy of Natural Sciences of Philadelphia 61: 194–234. -- Show included taxa

Banks, N. (1911). The pseudoscorpions of California. Journal of Entomology, Pomona College 3: 633–640. -- Show included taxa

Beier, M. (1932a). Pseudoscorpionidea I. Subord. Chthoniinea et Neobisiinea. Tierreich 57: i–xx, 1–258. -- Show included taxa

Beier, M. (1932g). Pseudoscorpionidea - Afterskorpione. In: Kükenthal, W. and Krumbach, T. (ed.) Handbuch der Zoologie Walter de Gruyter & Co., Berlin und Leipzig 3 (2) (5): 117–192. -- Show included taxa

Benavides, L.R., Cosgrove, J.G., Harvey, M.S. & Giribet, G. (2019). Phylogenomic interrogation resolves the backbone of the Pseudoscorpiones Tree of Life. Molecular Phylogenetics and Evolution 139(106509): 1–14. -- Show included taxa

Ceballos, A. (2004). Pseudoscorpionida. In: Bousquets, J.L., Morrone, J.J., Ordóñez, O.Y. and Fernández, I.V. (ed.) Biodiversidad, taxonomía y biogeografía de Artrópodos de México: hacia una síntesis de su conocimiento Facultad de Ciencias, UNAM, México, D.F. 4: 417–429. -- Show included taxa

Chamberlin, J.C. (1921). Notes on the genus Garypus in North America (Pseudoscorpionida - Cheliferidae). Canadian Entomologist 53: 186–191. -- Show included taxa

Chamberlin, J.C. (1925a). On a collection of pseudoscorpions from the stomach contents of toads. University of California Publications in Entomology 3: 327–332. -- Show included taxa

Chamberlin, J.C. (1930). A synoptic classification of the false scorpions or chela-spinners, with a report on a cosmopolitan collection of the same. Part II. The Diplosphyronida (Arachnida-Chelonethida). Annals and Magazine of Natural History (10) 5: 1–48, 585–620. -- Show included taxa

Chamberlin, J.C. (1931a). The arachnid order Chelonethida. Stanford University Publications, Biological Sciences 7(1): 1–284. -- Show included taxa

Chamberlin, J.C. (1935b). Chelonethida. In: Pratt, H.S. (ed.) A manual of the common invertebrate animals (exclusive of insects) Blakiston Company, Philadelphia and Toronto: 477–481. -- Show included taxa

Firstman, B. (1973). The relationship of the chelicerate arterial system to the evolution of the endosternite. Journal of Arachnology 1: 1–54. -- Show included taxa

Harvey, M.S. (1991a). Catalogue of the Pseudoscorpionida. Manchester University Press, Manchester. -- Show included taxa

Hoff, C.C. (1958). List of the pseudoscorpions of North America north of Mexico. American Museum Novitates 1875: 1–50. -- Show included taxa

Lee, V.F. (1979a). The maritime pseudoscorpions of Baja California, México (Arachnida: Pseudoscorpionida). Occasional Papers of the California Academy of Sciences 131: i–iv, 1–38. -- Show included taxa

Lee, V.F. (1979b). Unusual habitats of some Garypus pseudoscorpions of Baja California, México. American Arachnology 20: 13. -- Show included taxa

Lighton, J.R.B. & Joos, B. (2002a). Discontinuous gas exchange in the pseudoscorpion Garypus californicus is regulated by hypoxia, not hypercapnia. Physiological and Biochemical Zoology 75: 345–349. -- Show included taxa

Lighton, J.R.B. & Joos, B. (2002b). Discontinuous gas exchange in a tracheate arthropod, the pseudoscorpion Garypus californicus: occurrence, characteristics and temperature dependence. Journal of Insect Science 2(23): 1–4. -- Show included taxa

Moles, M. & Moore, W. (1921). A list of California Arachnida. I. Pseudoscorpionida. Journal of Entomology and Zoology, Pomona College 13: 6–9. -- Show included taxa

Moore, W.T. (1917). Record of two pseudoscorpions from Claremont-Laguna region. Journal of Entomology and Zoology, Pomona College 9: 26–29. -- Show included taxa

Muchmore, W.B. (1990a). Pseudoscorpionida. In: Dindal, D.L. (ed.) Soil biology guide John Wiley and Sons, New York: 503–527. -- Show included taxa

Newell, I.M. (1975). Marine mites (Halacaridae). Order Pseudoscorpionida (Chelonethida). Marine tardigrades. In: Smith, R.I. and Carlton, J.T. (ed.) Light's manual: intertidal invertebrates of the central California coast University of California Press, Berkeley: 425–432, 435. -- Show included taxa

Poinar, G.O., Jr, Thomas, G.M. & Lee, V.F. (1985). Laboratory infection of Garypus californicus (Pseudoscorpionida, Garypidae) with neoaplectanid and heterorhabditid nematodes (Rhabditoidea). Journal of Arachnology 13: 400–401. -- Show included taxa

Ricketts, E.F. & Calvin, J. (1960). Between Pacific tides. Stanford University Press, Stanford. [third edition (revised by J.W. Hedgpeth)] -- Show included taxa

Roewer, C.F. (1937). Chelonethi oder Pseudoskorpione. In: Bronns, H.G. (ed.) Bronn's Klassen und Ordnungen des Tierreichs Akademische Verlagsgesellschaft M.B.H., Leipzig 5(IV)(6)(1): 161–320. -- Show included taxa

Roth, V.D. & Brown, W.L. (1976). Other intertidal air-breathing arthropods. In: Cheng, L. (ed.) Marine insects North-Holland Publishing Co., Amsterdam and Oxford: 119–150. -- Show included taxa

Schulte, G. (1976). Litoralzonierung von Pseudoskorpionen an der nordamerikanischen Pazifikküste (Arachnida: Pseudoscorpiones: Neobisiidae, Garypidae). Entomologica Germanica 3: 119–124. -- Show included taxa

Zeh, D.W. (1987b). Aggression, density, and sexual dimorphism in chernetid pseudoscorpions (Arachnida: Pseudoscorpionida). Evolution 41: 1072–1087. -- Show included taxa

Harvey, M.S., Hillyer, M.J., Carvajal, J.I. & Huey, J.A. (2020). Supralittoral pseudoscorpions of the genus Garypus (Pseudoscorpiones: Garypidae) from the Indo-West Pacific region, with a review of the subfamily classification of Garypidae. Invertebrate Systematics 34: 34–87. -- Show included taxa

Date Type Reference Old value
2022-07-03 New taxonomic reference entry Benavides, Cosgrove, Harvey and Giribet, 2019
2021-12-23 Species updated Old value
2021-12-23 Species updated Old value
2021-12-23 New taxonomic reference entry Harvey, Hillyer, Carvajal and Huey, 2020