Pseudogarypinus frontalis (Banks, 1909)


Original combination: Olpium frontalis


Taxonomic references

Olpium frontalis

Banks, 1909b: 307 original description

Serianus (?) frontalis (Banks):

Beier, 1932a: 213
Roewer, 1937: 266

Pseudogarypinus frontalis (Banks):

Hoff, 1956b: 33–34
Benedict and Malcolm, 1978a: 125–132, figs 15–18
Muchmore, 1990a: 515, figs 18.43, 18.49
Harvey, 1991a: 299–300
Francke and Villegas-Guzmán, 2006: 295
Turienzo, Iorio and Mahnert, 2010: 561, 590
Benavides, Cosgrove, Harvey and Giribet, 2019: 3, 4, figs 3, 4

Pseudogarypinus (?) frontalis (Banks):

Hoff, 1958: 16–17
Hoff, 1959b: 4, etc

Garypinus marianae Chamberlin, 1930 -- synonymised by Benedict and Malcolm, 1978a: 125
Garypinus marianae
Chamberlin, 1930: 591–592, figs 1t, 1aa (synonymised by Benedict and Malcolm, 1978a: 125) original description
Chamberlin, 1931a: n/a, figs 11w, 15l, 19i, 26b, 37b, 47j
Pseudogarypinus marianae (Chamberlin):
Beier, 1931c: 314 (as Pseudogarypinus marinae [sic])
Beier, 1932a: 206–207
Roewer, 1937: 265
Gering, 1956: 50
Hoff, 1958: 16
Hoff, 1961: 439–440
Weygoldt, 1969a: 43, 124
Weygoldt, 1970a: 242, fig. 3
Muchmore, 1971a: 91
Knowlton, 1974: 3
Rowland and Reddell, 1976: 16
Schaller, 1979: n/a, plate 10.III fig. 3
Zeh, 1987b: 1086
Weygoldt, 1990: n/a, fig. 7A3
Distribution table

Realm: Nearctic

US (California, Colorado, New Mexico, Oregon, Texas, Utah, Washington)
Type locality: Las Vegas, San Miguel County, New Mexico, U.S.A.


Locality Lat Lon
1 Madrone Picnic Area 37.254444 -122.382777
Type repository
  • Lectotype female (MCZ). Paralectotype: 1 tritonymph (MCZ)
Zoobank LSID


Banks, N. (1909b). Arachnida from Costa Rica. Proceedings of the Academy of Natural Sciences of Philadelphia 61: 194–234. -- Show included taxa

Beier, M. (1931c). Neue Pseudoscorpione der U. O. Neobisiinea. Mitteilung aus dem Zoologischen Museum in Berlin 17: 299–318. -- Show included taxa

Beier, M. (1932a). Pseudoscorpionidea I. Subord. Chthoniinea et Neobisiinea. Tierreich 57: i–xx, 1–258. -- Show included taxa

Benavides, L.R., Cosgrove, J.G., Harvey, M.S. & Giribet, G. (2019). Phylogenomic interrogation resolves the backbone of the Pseudoscorpiones Tree of Life. Molecular Phylogenetics and Evolution 139(106509): 1–14. -- Show included taxa

Benedict, E.M. & Malcolm, D.R. (1978a). Some garypoid false scorpions from western North America (Pseudoscorpionida: Garypidae and Olpiidae). Journal of Arachnology 5: 113–132. -- Show included taxa

Chamberlin, J.C. (1930). A synoptic classification of the false scorpions or chela-spinners, with a report on a cosmopolitan collection of the same. Part II. The Diplosphyronida (Arachnida-Chelonethida). Annals and Magazine of Natural History (10) 5: 1–48, 585–620. -- Show included taxa

Chamberlin, J.C. (1931a). The arachnid order Chelonethida. Stanford University Publications, Biological Sciences 7(1): 1–284. -- Show included taxa

Francke, O.F. & Villegas-Guzmán, G.A. (2006). Symbiotic relationships between pseudoscorpions (Arachnida) and packrats (Rodentia). Journal of Arachnology 34: 289–298. -- Show included taxa

Gering, R.L. (1956). Arachnids: spiders, pseudoscorpions, scorpions, solpugids. In: Woodbury, A.M. (ed.) Ecological check lists. The Great Salt Lake Desert series University of Utah, Dugway. -- Show included taxa

Harvey, M.S. (1991a). Catalogue of the Pseudoscorpionida. Manchester University Press, Manchester. -- Show included taxa

Hoff, C.C. (1956b). Diplosphyronid pseudoscorpions from New Mexico. American Museum Novitates 1780: 1–49. -- Show included taxa

Hoff, C.C. (1958). List of the pseudoscorpions of North America north of Mexico. American Museum Novitates 1875: 1–50. -- Show included taxa

Hoff, C.C. (1959b). The ecology and distribution of the pseudoscorpions of north-central New Mexico. University of New Mexico Publications in Biology 8: 1–68. -- Show included taxa

Hoff, C.C. (1961). Pseudoscorpions from Colorado. Bulletin of the American Museum of Natural History 122: 409–464. -- Show included taxa

Knowlton, G.F. (1974). Some pseudoscorpions of Curlew Valley. Utah State University Ecology Center, Terrestrial Arthropods Series 11: 1–3. -- Show included taxa

Muchmore, W.B. (1971a). Phoresy by North and Central American pseudoscorpions. Proceedings of the Rochester Academy of Science 12: 79–97. -- Show included taxa

Muchmore, W.B. (1990a). Pseudoscorpionida. In: Dindal, D.L. (ed.) Soil biology guide John Wiley and Sons, New York: 503–527. -- Show included taxa

Roewer, C.F. (1937). Chelonethi oder Pseudoskorpione. In: Bronns, H.G. (ed.) Bronn's Klassen und Ordnungen des Tierreichs Akademische Verlagsgesellschaft M.B.H., Leipzig 5(IV)(6)(1): 161–320. -- Show included taxa

Rowland, J.M. & Reddell, J.R. (1976). Annotated checklist of the arachnid fauna of Texas (excluding Acarida and Araneida). Occasional Papers of the Museum, Texas Technical University 38: 1–25. -- Show included taxa

Schaller, F. (1979). Significance of sperm transfer and formation of spermatophores in arthropod phylogeny. In: Gupta, A.P. (ed.) Arthropod phylogeny Van Nostrand Reinhold, New York: 587–608. -- Show included taxa

Turienzo, P., Iorio, O. di & Mahnert, V. (2010). Global checklist of pseudoscorpions (Arachnida) found in birds’ nests. Revue Suisse de Zoologie 117: 557–598. -- Show included taxa

Weygoldt, P. (1969a). The biology of pseudoscorpions. Harvard University Press, Cambridge, Massachusetts. -- Show included taxa

Weygoldt, P. (1970a). Vergleichende Untersuchungen zur Fortpflanzungsbiologie der Pseudoscorpione II. Zeitschrift für die Zoologische Systematik und Evolutionforschung 8: 241–259. -- Show included taxa

Weygoldt, P. (1990). Arthropoda-Chelicerata: sperm transfer. In: Adiyodi, K.G. and Adiyodi, R.G. (ed.) Reproductive biology of invertebrates Oxford & IBH Publishing Co., New Delhi 4 (B): 77–119. -- Show included taxa

Zeh, D.W. (1987b). Aggression, density, and sexual dimorphism in chernetid pseudoscorpions (Arachnida: Pseudoscorpionida). Evolution 41: 1072–1087. -- Show included taxa

Date Type Reference Old value
2022-07-03 New taxonomic reference entry Benavides, Cosgrove, Harvey and Giribet, 2019