Taxa included

Morikawa, K. (1960). Systematic studies of Japanese pseudoscorpions. Memoirs of Ehime University (2B) 4: 85–172.

Taxon
Pseudoscorpiones de Geer, 1778 92, Chthoniinea
Chthoniidae Daday, 1889 (Pseudoscorpiones) 92–93, Dithidae Chamberlin
Chthoniidae Daday, 1889 (Pseudoscorpiones) 93, Chthoniinae
Ditha (Paraditha) marcusensis (Morikawa, 1952) (Pseudoscorpiones: Chthoniidae) 93, plate 1 fig. 1, plate 7 fig. 1, Ditha marcusensis (Morikawa)
Ditha Chamberlin, 1929 (Pseudoscorpiones: Chthoniidae) 93, Ditha
Chthoniidae Daday, 1889 (Pseudoscorpiones) 93, Chthoniidae Daday
Mundochthonius japonicus Chamberlin, 1929 (Pseudoscorpiones: Chthoniidae) 94–95, Mundochthonius japonicus
Mundochthonius Chamberlin, 1929 (Pseudoscorpiones: Chthoniidae) 94, Mundochthonius
Mundochthonius japonicus scolytidis Morikawa, 1954 (Pseudoscorpiones: Chthoniidae) 95–96, plate 1 fig. 2, plate 10 fig. 9, Mundochthonius japonicus scolyditis Morikawa
Mundochthonius japonicus japonicus Chamberlin, 1929 (Pseudoscorpiones: Chthoniidae) 95, plate 7 fig. 2, Mundochthonius japonicus japonicus Chamberlin
Mundochthonius japonicus tripartitus Morikawa, 1956 (Pseudoscorpiones: Chthoniidae) 96, plate 10 fig. 6, Mundochthonius japonicus tripartitus Morikawa
Mundochthonius japonicus imadatei Morikawa, 1956 (Pseudoscorpiones: Chthoniidae) 96–97, Mundochthonius japonicus imadatei Morikawa
Allochthonius Chamberlin, 1929 (Pseudoscorpiones: Pseudotyrannochthoniidae) 97, Allochthonius
Allochthonius (Allochthonius) opticus (Ellingsen, 1907) (Pseudoscorpiones: Pseudotyrannochthoniidae) 98, Allochthonius (Allochthonius) opticus (Ellingsen)
Allochthonius (Allochthonius) opticus opticus (Ellingsen, 1907) (Pseudoscorpiones: Pseudotyrannochthoniidae) 99–100, plate 1 figs 3–4, plate 5 fig. 1, plate 9 fig. 1, plate 10 figs 10–12, Allochthonius (Allochthonius) opticus opticus (Ellingsen)
Allochthonius (Allochthonius) opticus troglophilus Morikawa, 1956 (Pseudoscorpiones: Pseudotyrannochthoniidae) 100–101, plate 5 fig. 2, Allochthonius (Allochthonius) opticus troglophilus Morikawa
Allochthonius (Urochthonius) biocularis Morikawa, 1956 (Pseudoscorpiones: Pseudotyrannochthoniidae) 101, plate 5 fig. 7, Allochthonius (Urochthonius) biocularis
Allochthonius (Urochthonius) ishikawai Morikawa, 1954 (Pseudoscorpiones: Pseudotyrannochthoniidae) 101–102, Allochthonius (Urochthonius) ishikawai
Allochthonius (Urochthonius) ishikawai shiragatakiensis Morikawa, 1954 (Pseudoscorpiones: Pseudotyrannochthoniidae) 102–103, plate 7 fig. 5, Allochthonius (Urochthonius) ishikawai shiragatakiensis
Allochthonius (Urochthonius) ishikawai ishikawai Morikawa, 1954 (Pseudoscorpiones: Pseudotyrannochthoniidae) 102, plate 1 fig. 6, plate 7 fig. 3, plate 5 fig. 5, plate 8 fig. 12, Allochthonius (Urochthonius) ishikawai ishikawai Morikawa
Allochthonius (Urochthonius) ishikawai kyushuensis Morikawa, 1960 (Pseudoscorpiones: Pseudotyrannochthoniidae) 103–104, plate 1 fig. 5, plate 4 fig. 12, plate 5 fig. 9, Allochthonius (Urochthonius) ishikawai kyushuensis
Allochthonius (Urochthonius) ishikawai uenoi Morikawa, 1956 (Pseudoscorpiones: Pseudotyrannochthoniidae) 103, plate 5 fig. 3, plate 10 fig. 5, Allochthonius (Urochthonius) ishikawai uenoi Morikawa
Allochthonius (Urochthonius) ishikawai uyamadensis Morikawa, 1954 (Pseudoscorpiones: Pseudotyrannochthoniidae) 103, plate 7 fig. 4, plate 5 figs 8, 11, Allochthonius (Urochthonius) ishikawai uyamadensis Morikawa
Allochthonius (Urochthonius) ishikawai deciclavatus Morikawa, 1956 (Pseudoscorpiones: Pseudotyrannochthoniidae) 104, plate 5 fig. 6, Allochthonius (Urochthonius) ishikawai deciclavatus Morikawa
Spelaeochthonius Morikawa, 1954 (Pseudoscorpiones: Pseudotyrannochthoniidae) 104–105, Allochthonius Morikawa
Pseudotyrannochthonius undecimclavatus (Morikawa, 1956) (Pseudoscorpiones: Pseudotyrannochthoniidae) 105, Allochthonius (Spelaeochthonius) undecimclavatus (Morikawa)
Pseudotyrannochthonius undecimclavatus undecimclavatus (Morikawa, 1956) (Pseudoscorpiones: Pseudotyrannochthoniidae) 106, plate 5 figs 10, 14, plate 10 fig. 4, Allochthonius (Spelaeochthonius) undecimclavatus undecimclavatus (Morikawa)
Pseudotyrannochthonius undecimclavatus kishidai (Morikawa, 1960) (Pseudoscorpiones: Pseudotyrannochthoniidae) 106–107, plate 5 fig. 16, Allochthonius (Spelaeochthonius) undecimclavatus kishidai
Pseudotyrannochthonius kobayashii kobayashii (Morikawa, 1956) (Pseudoscorpiones: Pseudotyrannochthoniidae) 107–108, plate 5 fig. 13, plate 10 fig. 1, Allochthonius (Spelaeochthonius) kobayashii kobayashii (Morikawa)
Pseudotyrannochthonius kobayashii (Morikawa, 1956) (Pseudoscorpiones: Pseudotyrannochthoniidae) 107, Allochthonius (Spelaeochthonius) kobayashii Morikawa
Tyrannochthonius Chamberlin, 1929 (Pseudoscorpiones: Chthoniidae) 108, Tyrannochthonius
Pseudotyrannochthonius kubotai (Morikawa, 1954) (Pseudoscorpiones: Pseudotyrannochthoniidae) 108, plate 1 fig. 8, plate 5 fig. 12, plate 9 fig. 2, plate 10 fig. 3, Allochthonius (Spelaeochthonius) kubotai (Morikawa)
Pseudotyrannochthonius kobayashii dorogawaensis (Morikawa, 1956) (Pseudoscorpiones: Pseudotyrannochthoniidae) 108, plate 1 fig. 7, plate 10 fig. 2, Allochthonius (Spelaeochthonius) kobayashii dorogawaensis (Morikawa)
Pseudotyrannochthonius kobayashii akiyoshiensis (Morikawa, 1956) (Pseudoscorpiones: Pseudotyrannochthoniidae) 108, plate 6 fig. 6, plate 5 fig. 15, Allochthonius (Spelaeochthonius) kobayashii akiyoshiensis (Morikawa)
Tyrannochthonius japonicus (Ellingsen, 1907) (Pseudoscorpiones: Chthoniidae) 109, Tyrannochthonius japonicus (Ellingsen)
Tyrannochthonius japonicus japonicus (Ellingsen, 1907) (Pseudoscorpiones: Chthoniidae) 109–110, Tyrannochthonius japonicus japonicus (Ellingsen)
Paraliochthonius takashimai (Morikawa, 1958) (Pseudoscorpiones: Chthoniidae) 110, plate 5 fig. 4, plate 10 fig. 8, Tyrannochthonius takashimai Morikawa
Tyrannochthonius japonicus dogoensis Morikawa, 1954 (Pseudoscorpiones: Chthoniidae) 110, plate 1 fig. 9, plate 7 fig. 9, plate 10 fig. 7, Tyrannochthonius japonicus dogoensis Morikawa
Lechytia Balzan, 1892 (Pseudoscorpiones: Chthoniidae) 111, Lechytia
Pseudoscorpiones de Geer, 1778 111, Neobisiinea
Chthoniidae Daday, 1889 (Pseudoscorpiones) 111, Lechytiinae Chamberlin
Lechytia sakagamii Morikawa, 1952 (Pseudoscorpiones: Chthoniidae) 111, plate 2 fig. 1, plate 7 fig. 10, Lechytia sakagamii
Neobisium Chamberlin, 1930 (Pseudoscorpiones: Neobisiidae) 112, Neobisium
Neobisiidae Chamberlin, 1930 (Pseudoscorpiones) 112, Neobisiidae
Parobisium Chamberlin, 1930 (Pseudoscorpiones: Neobisiidae) 112–113, Neobisium
Parobisium magnum ohuyeanum (Morikawa, 1952) (Pseudoscorpiones: Neobisiidae) 113–114, plate 2 fig. 2, plate 6 fig. 4, plate 9 fig. 3, Neobisium (Parobisium) magnum ohuyeanum Morikawa
Parobisium magnum magnum (Chamberlin, 1930) (Pseudoscorpiones: Neobisiidae) 113, (in part, see Parobisium imperfectum (Chamberlin)), Neobisium (Parobisium) magnum magnum Chamberlin
Parobisium magnum (Chamberlin, 1930) (Pseudoscorpiones: Neobisiidae) 113, Neobisium (Parobisium) magnum
Parobisium imperfectum (Chamberlin, 1930) (Pseudoscorpiones: Neobisiidae) 113, (in part), Neobisium (Parobisium) magnum magnum Chamberlin
Microbisium pygmaeum (Ellingsen, 1907) (Pseudoscorpiones: Neobisiidae) 114–115, plate 2 fig. 3, Neobisium (Parobisium) pygmaeum (Ellingsen)
Parobisium flexifemoratum (Chamberlin, 1930) (Pseudoscorpiones: Neobisiidae) 114, Neobisium (Parobisium) flexifemoratum
Parobisium anagamidense anagamidense (Morikawa, 1957) (Pseudoscorpiones: Neobisiidae) 115, plate 6 fig. 1, plate 9 fig. 4 (as Neobisium (Parobisium) anagamidensis anagamidensis), Neobisium (Parobisium) anagamidense anagamidense Morikawa
Parobisium anagamidense (Morikawa, 1957) (Pseudoscorpiones: Neobisiidae) 115, (as Neobisium (Parobisium) anagamidensis), Neobisium (Parobisium) anagamidense
Neobisium (Parobisium) anagamidense longidigitatum Morikawa, 1957 (Pseudoscorpiones: Neobisiidae) 115–116, (as Neobisium (Parobisium) anagamidensis longidigitatus [sic]), Neobisium (Parobisium) anagamidense longidigitatum
Parobisium anagamidense esakii (Morikawa, 1960) (Pseudoscorpiones: Neobisiidae) 116, (as Neobisium (Parobisium) anagamidensis esakii), Neobisium (Parobisium) anagamidense esakii
Roncus L. Koch, 1873 (Pseudoscorpiones: Neobisiidae) 116, Roncus
Pararoncus japonicus (Ellingsen, 1907) (Pseudoscorpiones: Syarinidae) 117–118, plate 2 fig. 4, plate 7 fig. 11 (in part; see Pararoncus histrionicus Chamberlin), Roncus (Roncus) japonicus (Ellingsen)
Pararoncus histrionicus Chamberlin, 1938 (Pseudoscorpiones: Syarinidae) 117–118, (misidentification, in part), Roncus (Roncus) japonicus (Ellingsen)
Pararoncus chamberlini (Morikawa, 1960) (Pseudoscorpiones: Syarinidae) 117–118, Roncus (Roncus) chamberlini
Pararoncus yosii (Morikawa, 1960) (Pseudoscorpiones: Syarinidae) 118, Roncus (Roncus) chamberlini yosii
Roncus (Roncus) troglophilus (Morikawa, 1957) (Pseudoscorpiones: Neobisiidae) 118, plate 6 figs 2–3, plate 9 figs 7, 9, Roncus (Roncus) chamberlini chamberlini Morikawa
Pararoncus oinuanensis (Morikawa, 1957) (Pseudoscorpiones: Syarinidae) 118, Roncus (Roncus) chamberlini oinuanensis Morikawa
Pararoncus chamberlini (Morikawa, 1960) (Pseudoscorpiones: Syarinidae) 118, plate 6 figs 2–3, plate 9 figs 7, 9, Roncus (Roncus) chamberlini chamberlini Morikawa
Pararoncus uenoi (Morikawa, 1957) (Pseudoscorpiones: Syarinidae) 119, plate 6 fig. 9, plate 9 fig. 6, Roncus (Parablothrus) uenoi Morikawa
Pararoncus rakanensis (Morikawa, 1957) (Pseudoscorpiones: Syarinidae) 119, plate 2 fig. 5, plate 6 figs 5–6, plate 9 fig. 5, Roncus (Parablothrus) rakanensis Morikawa
Halobisium Chamberlin, 1930 (Pseudoscorpiones: Neobisiidae) 119, Halobisium
Halobisium orientale japonicum Morikawa, 1958 (Pseudoscorpiones: Neobisiidae) 119–120, plate 2 fig. 6, plate 6 fig. 7, Halobisium orientale japonicum
Microcreagris Balzan, 1892 (Pseudoscorpiones: Neobisiidae) 120, Microcreagris
Bisetocreagris brevidigitata (Chamberlin, 1930) (Pseudoscorpiones: Neobisiidae) 121, Microcreagris brevidigitata
Bisetocreagris japonica (Ellingsen, 1907) (Pseudoscorpiones: Neobisiidae) 121–123, plate 2 fig. 7, plate 6 fig. 11, plate 8 fig. 7, plate 9 fig. 8, plate 10 fig. 21, Microcreagris japonica Ellingsen
Bisetocreagris pygmaea (Ellingsen, 1907) (Pseudoscorpiones: Neobisiidae) 123, Microcreagris pygmaea
Microcreagris microdivergens Morikawa, 1955 (Pseudoscorpiones: Neobisiidae) 123, Microcreagris microdivergens
Microcreagris macropalpus Morikawa, 1955 (Pseudoscorpiones: Neobisiidae) 123, plate 4 fig. 9, plate 8 fig. 8, plate 9 fig. 10, plate 10 fig. 22, Microcreagris macropalpus Morikawa
Microcreagris microdivergens Morikawa, 1955 (Pseudoscorpiones: Neobisiidae) 124, plate 4 fig. 11, plate 6 fig. 10, plate 8 fig. 9, plate 9 fig. 11, plate 10 fig. 23, Microcreagris microdivergens microdivergens Morikawa
Microcreagris pseudoformosa Morikawa, 1955 (Pseudoscorpiones: Neobisiidae) 124, plate 4 fig. 10, plate 8 fig. 11, plate 9 fig. 14, plate 10 fig. 26, Microcreagris pseudoformosa
Olpiidae Banks, 1895 (Pseudoscorpiones) 125, Olpiidae Banks
Xenolpium Chamberlin, 1930 (Pseudoscorpiones: Olpiidae) 126, Xenolpium
Nipponogarypus Morikawa, 1955 (Pseudoscorpiones: Olpiidae) 126–127, Nipponogarypus
Xenolpium oceanicum palauense Beier, 1957 (Pseudoscorpiones: Olpiidae) 126, Xenolpium oceanicum palauense
Nipponogarypus enoshimaensis enoshimaensis Morikawa, 1955 (Pseudoscorpiones: Olpiidae) 127–128, plate 2 fig. 8, plate 6 fig. 8, plate 7 fig. 14, plate 10 figs 32–33, Nipponogarypus enoshimaensis enoshimaensis Morikawa
Nipponogarypus enoshimaensis Morikawa, 1955 (Pseudoscorpiones: Olpiidae) 127, Nipponogarypus enoshimaensis
Nipponogarypus enoshimaensis okinoerabensis Morikawa, 1960 (Pseudoscorpiones: Olpiidae) 128, Nipponogarypus enoshimaensis okineroabensis
Garypinidae Daday, 1889 (Pseudoscorpiones) 129, Garypininae
Amblyolpium japonicum Morikawa, 1960 (Pseudoscorpiones: Garypinidae) 129–130, plate 2 fig. 9, plate 7 fig. 12, plate 9 fig. 15, Amblyolpium japonicum
Amblyolpium Simon, 1898 (Pseudoscorpiones: Garypinidae) 129, Amblyolpium
Solinus Chamberlin, 1930 (Pseudoscorpiones: Garypinidae) 130, Solinus
Garypidae Simon, 1879 (Pseudoscorpiones) 130–131, Garypidae Simon
Solinus japonicus Morikawa, 1953 (Pseudoscorpiones: Garypinidae) 130, plate 3 fig. 3, plate 7 fig. 13, Solinus japonicus
Garypus L. Koch, 1873 (Pseudoscorpiones: Garypidae) 131, Garypus
Garypidae Simon, 1879 (Pseudoscorpiones) 131, Garypinae
Anchigarypus japonicus (Beier, 1952) (Pseudoscorpiones: Garypidae) 131–132, plate 3 fig. 1, plate 4 figs 7–8, plate 6 fig. 12, plate 10 fig. 19, Garypus japonicus
Geogarypus (Geogarypus) micronesiensis Morikawa, 1952 (Pseudoscorpiones: Geogarypidae) 132–133, plate 3 fig. 2, plate 7 fig. 15, Geogarypus (Geogarypus) micronesiensis
Geogarypus Chamberlin, 1930 (Pseudoscorpiones: Geogarypidae) 132, Geogarypus Chamberlin
Geogarypus Chamberlin, 1930 (Pseudoscorpiones: Geogarypidae) 132, Geogarypus
Geogarypidae Chamberlin, 1930 (Pseudoscorpiones) 132, Geogarypinae
Cheiridiidae Hansen, 1894 (Pseudoscorpiones) 133–134, Cheiridiidae Hansen
Pseudoscorpiones de Geer, 1778 133, Cheliferinea
Cheiridium minor Chamberlin, 1938 (Pseudoscorpiones: Cheiridiidae) 134–135, plate 3 fig. 4, plate 10 figs 18, 27, Cheiridium minor Chamberlin
Cheiridium Menge, 1855 (Pseudoscorpiones: Cheiridiidae) 134, Cheiridium
Apocheiridium pinium Morikawa, 1953 (Pseudoscorpiones: Cheiridiidae) 135, plate 3 fig. 5, plate 8 fig. 15, plate 9 fig. 16, plate 10 fig. 20, Apocheiridium pinium
Apocheiridium Chamberlin, 1924 (Pseudoscorpiones: Cheiridiidae) 135, Apocheiridium
Atemnidae Kishida, 1929 (Pseudoscorpiones) 136, Atemnidae
Paratemnus Beier, 1932 (Pseudoscorpiones: Atemnidae) 137, Paratemnus
Oratemnus samoanus whartoni Chamberlin, 1947 (Pseudoscorpiones: Atemnidae) 138–139, Oratemnus samoanus whartoni Chamberlin
Paratemnoides japonicus (Morikawa, 1953) (Pseudoscorpiones: Atemnidae) 138, plate 3 fig. 6, plate 8 figs 1, 16, plate 9 fig. 17, Paratemnus japonicus
Paratemnoides philippinus (Beier, 1932) (Pseudoscorpiones: Atemnidae) 138, Paratemnus philippinus
Paratemnoides philippinus (Beier, 1932) (Pseudoscorpiones: Atemnidae) 138, Paratemnus philippinus
Oratemnus Beier, 1932 (Pseudoscorpiones: Atemnidae) 138, Oratemnus
Chernetidae Menge, 1855 (Pseudoscorpiones) 139, Chernetidae
Muscichernes katoi Morikawa, 1960 (Pseudoscorpiones: Chernetidae) 140–141, plate 3 fig. 7, plate 7 fig. 16, plate 9 fig. 19, Muscichernes katoi
Muscichernes Morikawa, 1960 (Pseudoscorpiones: Chernetidae) 140, (synonymised with Lamprochernes by Harvey, 1987c: 111), Muscichernes
Haplochernes Beier, 1932 (Pseudoscorpiones: Chernetidae) 143, Haplochernes
Haplochernes boncicus boncicus (Karsch, 1881) (Pseudoscorpiones: Chernetidae) 143, Haplochernes boncicus boncicus (Karsch)
Haplochernes boncicus (Karsch, 1881) (Pseudoscorpiones: Chernetidae) 143, Haplochernes boncicus (Karsch)
Haplochernes boncicus hagai Morikawa, 1953 (Pseudoscorpiones: Chernetidae) 144, plate 3 fig. 8, plate 8 figs 2, 14, plate 9 fig. 22, Haplochernes boncicus hagai Morikawa
Megachernes Beier, 1932 (Pseudoscorpiones: Chernetidae) 144, Megachernes
Megachernes ryugadensis Morikawa, 1954 (Pseudoscorpiones: Chernetidae) 144–145, Megachernes ryugadensis
Haplochernes boninensis Beier, 1957 (Pseudoscorpiones: Chernetidae) 144, Haplochernes boninensis
Megachernes ryugadensis ryugadensis Morikawa, 1954 (Pseudoscorpiones: Chernetidae) 145, plate 7 fig. 17, plate 10 fig. 28, Megachernes ryugadensis ryugadensis Morikawa
Megachernes ryugadensis naikaiensis Morikawa, 1957 (Pseudoscorpiones: Chernetidae) 145–146, plate 3 fig. 9, plate 9 fig. 23, Megachernes ryugadensis naikaiensis
Allochernes Beier, 1932 (Pseudoscorpiones: Chernetidae) 146, Allochernes
Megachernes ryugadensis myophilus Morikawa, 1960 (Pseudoscorpiones: Chernetidae) 146, Megachernes ryugadensis myophilus
Allochernes japonicus (Morikawa, 1953) (Pseudoscorpiones: Chernetidae) 147, plate 4 fig. 1, plate 8 fig. 3, plate 9 fig. 21, Allochernes (Toxochernes) japonicus (Morikawa)
Allochernes ginkgoanus (Morikawa, 1953) (Pseudoscorpiones: Chernetidae) 147, plate 4 fig. 2, plate 8 fig. 5, plate 9 fig. 24, plate 10 fig. 17, Allochernes (Toxochernes) ginkgoanus (Morikawa)
Cheliferidae Risso, 1827 (Pseudoscorpiones) 148, Cheliferidae
Withius Kew, 1911 (Pseudoscorpiones: Withiidae) 149, Withius Kew
Withius japonicus Morikawa, 1954 (Pseudoscorpiones: Withiidae) 149, plate 4 fig. 3, plate 7 fig. 19, plate 8 fig. 13, plate 9 fig. 25, plate 10 figs 16, 29, Withius japonicus
Chelifer Geoffroy, 1762 (Pseudoscorpiones: Cheliferidae) 150–151, Chelifer
Kashimachelifer cinnamomeus Morikawa, 1957 (Pseudoscorpiones: Cheliferidae) 151–152, plate 4 fig. 4, plate 7 fig. 18, plate 6 fig. 13, plate 8 fig. 17, plate 9 fig. 27, plate 10 figs 15, 31, Kashimachelifer cinnamomeus
Kashimachelifer Morikawa, 1957 (Pseudoscorpiones: Cheliferidae) 151, Kashimachelifer
Chelifer cancroides orientalis Morikawa, 1954 (Pseudoscorpiones: Cheliferidae) 151, plate 4 fig. 5, plate 7 fig. 20, plate 8 fig. 18, plate 9 fig. 26, plate 10 fig. 23, Chelifer cancroides orientalis
Lophochernes bicarinatus Simon, 1878 (Pseudoscorpiones: Cheliferidae) 152–153, plate 4 fig. 6, plate 8 figs 4, 19, plate 9 fig. 28, plate 10 figs 14, 30, Lophochernes bicarinatus
Lophochernes sauteri (Ellingsen, 1907) (Pseudoscorpiones: Cheliferidae) 152, Lophochernes sauteri (Ellingsen)
Lophochernes Simon, 1878 (Pseudoscorpiones: Cheliferidae) 152, Lophochernes