Chthonius orthodactylus (Leach, 1817)

SPECIES |   ACCEPTED

Original combination: Obisium orthodactylum

LSID urn:lsid:wac.nmbe.ch:name:0059cd2e-553a-48a1-8bed-d1dc127adb11


Genus type of Chthonius C.L. Koch, 1843

Taxonomic references

Obisium orthodactylum

Leach, 1817: 51, plate 141 fig. 2 original description
Lucas, 1840: 450
Tulk, 1844: 55–57

Obisium ischnocheles Hermann:

Théis, 1832: 63–66, plate 1 figs 3, 3a–c (misidentification)

Chthonius orthodactylus (Leach):

Koch, 1843: 79–80, fig. 808
Hagen, 1870: 268
Koch, 1873: 50–51
Canestrini, 1875a: 2‒3
Koch, 1877: 181
Simon, 1879a: 73–74
Tömösváry, 1883: 239–240, plate 5 figs 3–8
Becker, 1884: cclxiv
Pavesi, 1884: 457–458
Canestrini, 1885: n/a, no. 8, figs 1–4
Simon, 1885c: 50
Daday, 1889a: 135, 191–192
Daday, 1889d: 25
Daday, 1889e: 81
Pickard-Cambridge, 1892: 209, plate A figs 7–9
Becker, 1896: 335
Simon, 1898d: 22 (misidentification, in part; see Paraliochthonius singularis (Menozzi))
Carpenter and Evans, 1899: 180–181
Simon, 1900b: 595
Evans, 1901a: 54
Carpenter, 1902: 220
Godfrey, 1908: 160
Bignotti, 1909: 67
Ellingsen, 1909a: 211, 216
Warburton, 1909: 438
Ellingsen, 1910a: 401
Godfrey, 1910: 30
Jackson, 1910: 50
Kew, 1911a: 57, fig. 20
Kew, 1916a: 124
Daday, 1918: 2
Caporiacco, 1926: 75
Beier, 1928: 312–313
Kästner, 1928: 9
Schenkel, 1928: 71
Beier, 1929a: 365
Beier, 1929e: 454
Berland, 1929: 18
Bristowe, 1929a: 25
Chamberlin, 1929a: 71
Kew, 1930: 253–256, figs 1c–d
Vachon, 1934g: 163
Ionescu, 1936: 2
Roewer, 1937: 239
Savory, 1945: 180, 182
Caporiacco, 1950a: 116
Caporiacco, 1951b: 103
Franciscolo, 1955: 125
Cerruti, 1959: 51 (as Chtonius [sic] orthodactylus)
Baggini, 1961: 24
Bishop, 1967: 393
Marcuzzi, 1968: 267
Szalay, 1968: 35–36, fig. 23c
Lazzeroni, 1970a: 207
Verner, 1971: n/a, fig. 4
Howes, 1972a: 108
Salmon, 1972: 67
Legg, 1973b: 430
Palmgren, 1973: 8
Legg, 1975a: 66
Legg, 1975b: 101, figs 1c, 4, 7c, 13a–b
Goddard, 1976b: 298–302, figs 2, 4
Goddard, 1979a: 100–104
Goddard, 1979b: 32
Judson, 1979a: 61
Matsuda, 1979: 201
Rundle, 1979: 47
Jones, 1980c: n/a, map 4
Schawaller, 1981b: 44, 50
Judson, 1987: 368
El-Hennawy, 1988b: 9
Harvey, 1992: 150
Fowles, 1994: 17
Legg and O'Connor, 1997: 109, fig. 2
Bell, Gates, Haughton, Macdonald, Smith, Wheater and Cullen, 1999: 237–239
Ducháč, 1999a: 14
Kárpáthegyi, 2007b: 91
Cawley, 2016: 117

Chthonius (Chthonius) orthodactylus (Leach):

Beier, 1932a: 50–51, figs 51, 56
Bristowe, 1938: 212
Beier, 1939e: 305
Cooreman, 1946b: 3
Beier, 1952e: 2
Beier, 1953e: 105
Beier and Franz, 1954: 453–454
Evans and Browning, 1954: 10, figs 9, 12
Beier, 1958c: 137
Beier, 1961f: 123
Beier, 1963b: 30, fig. 22
Beier, 1963a: 256
Beier, 1963h: 150
Lazzeroni, 1969a: 323
Lazzeroni, 1969b: 393–394
Lazzeroni, 1969c: 224
Lazzeroni, 1970c: 102
Legg, 1971a: 475, figs 1d, 2e
Ćurčić, 1974a: 12
Gardini, 1975: 2
Costantini, 1976: 122
Ćurčić, 1976b: 172
Crocker, 1978: 8
Leclerc and Heurtault, 1979: 239
Callaini, 1980a: 226
Gardini, 1980c: 100, 120, 121
Krumpál, 1980: 23
Inzaghi, 1981: 72
Krumpál and Kiefer, 1981: 128–129
Callaini, 1983d: 221–222
Bologna and Taglianti, 1985: 224, fig. 8
Callaini, 1986e: 236–237
Legg and Jones, 1988: 71–72, figs 14a, 14b(a–f)
Ducháč, 1989a: 180
Gardini, 1991a: 172
Harvey, 1991a: 157
Ducháč, 1994: 140
Gardini, 1995a: 2
Platen, Blick, Bliss, Drogla, Malten, Martens, Sacher and Wunderlich, 1995: 54
Drogla and Blick, 1996: 37
Gardini, 2000: 26–28
Henderickx, 1999b: 67
Šťáhlavský, 2001: 80, fig. 2
Delfosse, 2003b: 29
Mahnert, 2004: 463
Šťáhlavský and Král, 2004: 51, figs 1c, 5c
Ćurčić, Dimitrijević and Legakis, 2004: 235
Ozimec, 2004: 385
Tooren, 2005b: 92
Groh, 2007: 205
Kárpáthegyi, 2007a: 82
Petrov and Šťáhlavský, 2007: 16
Šťáhlavský and Krásný, 2007: 430
Christophoryová, Štáhlavský and Fedor, 2011: 36, fig. 2A
Isaia, Paschetta, Lana, Pantini, Schönhofer, Christian and Badino, 2011: 207, fig. 5.4
Mahnert, 2011d: 34
Šťáhlavský, 2011: 252
Christophoryová, Štáhlavský, Krumpál and Fedor, 2012: 5
Krajčovičová, Christophoryová and Krumpál, 2013: 715–716
Mahnert and Weber, 2013: 109, fig. 3
Šťáhlavský and Chytil, 2013: 80

Not Obisium orthodactylum Leach:

McIntire, 1868: 8 (misidentification; see Ephippiochthonius tetrachelatus)
McIntire, 1869: 247, fig. 219 (misidentification; see Ephippiochthonius tetrachelatus)

Not Chthonius orthodactylus (Leach):

Morley, 1930: 140–141 (misidentification; see Chthonius (Chthonius) ischnocheles (Hermann)
Morley, 1940: 167 (misidentification; see Chthonius (Chthonius) ischnocheles (Hermann))

Chthonius orthodactylus (Leach):

Distribution
Distribution table
Austria
Belgium
Bosnia
Bulgaria
Croatia
Czechia
France (mainland, Corse)
Germany
Greece (mainland)
Hungary
Ireland
Italy (mainland, Sardinia, Sicily)
Luxembourg
Montenegro
Netherlands
Romania
Serbia
Slovakia
Slovenia
Tunisia
UK
Type locality: Devon (as Danmonia), England, United Kingdom. Kent (as Cantia), England, United Kingdom.

Coordinates

Locality Lat Lon
1 Devon (IMPRECISE) 50.75 -4
2 Kent (IMPRECISE) 51.2 0.73
Type repository
    N/A
Zoobank LSID

urn:lsid:zoobank.org:act:E3255A12-45BB-4CDD-B793-70F0DD15980A

References

Baggini, A. (1961). Sulla distribuzione della fauna cavernicola Italiana nelle categorie biospeleologiche. Rassegna Speleologica Italiana 13: 1–35. -- Show included taxa

Becker, L. (1884). Catalogue des Arachnides de Belgique, cinquième partie. Chernètes. Comptes Rendus des Séances de la Société Entomologique de Belgique (3) no. 49 3: cclxiii–cclxiv. -- Show included taxa

Becker, L. (1896). Les Arachnides de Belgique. Annales de la Musée Royal d'Histoire Naturelle de Belgique 12: 1–378. -- Show included taxa

Beier, M. (1928). Die Pseudoskorpione des Wiener Naturhistorischen Museums. I. Hemictenodactyli. Annalen des Naturhistorischen Museums in Wien 42: 285–314. -- Show included taxa

Beier, M. (1929a). Die Pseudoskorpione des Wiener Naturhistorischen Museums. II. Panctenodactyli. Annalen des Naturhistorischen Museums in Wien 43: 341–367. -- Show included taxa

Beier, M. (1929e). Zoologische Forschungsreise nach den Jonischen Inseln und dem Peloponnes. I. und II. Teil. Sitzungsberichte der Akademie der Wissenschaften in Wien 128: 425–456. -- Show included taxa

Beier, M. (1932a). Pseudoscorpionidea I. Subord. Chthoniinea et Neobisiinea. Tierreich 57: i–xx, 1–258. -- Show included taxa

Beier, M. (1939e). Die Pseudoscorpione des oberösterreichischen Landesmuseums in Linz. Jahrbuch des Vereines für Landeskunde und Heimatpflege im Gau Oberdonau, Linz 88: 303–312. -- Show included taxa

Beier, M. (1952e). Ordn.: Pseudoscorpionidea, Afterskorpione. In: Strouhal, H. (ed.) Catalogus faunae Austriae Springer-Verlag, Wien 9a: 2–6. -- Show included taxa

Beier, M. (1953e). Ueber eine Pseudoscorpioniden-Ausbeute aus ligurischen Höhlen. Bollettino della Societa Entomologica Italiana 83: 105–108. -- Show included taxa

Beier, M. (1958c). Eine neue Neobisium-Art aus den Picentinischen Bergen in Süd-Italien. Memorie del Museo Civico di Storia Naturale di Verona 6: 135–137. -- Show included taxa

Beier, M. (1961f). Höhlenpseudoscorpione aus der Toscana. Monitore Zoologico Italiano 68: 123–127. -- Show included taxa

Beier, M. (1963a). Sizilianische Pseudoscorpione. Bolletino Accademia Gioenia di Scienze Naturali in Catania (4) 7: 253–263. -- Show included taxa

Beier, M. (1963b). Ordnung Pseudoscorpionidea (Afterskorpione). In: Bestimmungsbücher zur Bodenfauna Europas Akademie-Verlag, Berlin 1: vi, 313 pp.. -- Show included taxa

Beier, M. (1963h). Pseudoscorpione aus dem Museum "Enrico Caffi" in Bergamo. Rendiconti dell'Instituto Lombardo di Scienze e Lettere 97B: 147–156. -- Show included taxa

Beier, M. & Franz, H. (1954). 16. Ordnung: Pseudoscorpionidea. In: Franz, H. (ed.) Die Nordost-Alpen im Spiegel ihrer Landtierwelt Universitätsverlag Wagner, Innsbruck 1: 453–459. -- Show included taxa

Bell, J.R., Gates, S., Haughton, A.J., Macdonald, D.W., Smith, H., Wheater, C.P. & Cullen, W.R. (1999). Pseudoscorpions in field margins: effects of margin age, management and boundary habitats. Journal of Arachnology 27: 236–240. -- Show included taxa

Berland, L. (1929). Arachnides et Crustacès. In: Perrier, R. (ed.) Faune de la France en tableaux synoptiques Librairie Delagrave, Paris 2: 1–220. -- Show included taxa

Bignotti, G. (1909). Elenco dei Pseudoscorpioni trovati in Italia e loro distribuzione geografica. Atti della Società dei Naturaliste e Matematici, Modena (4) 11: 56–76. -- Show included taxa

Bishop, C.P. (1967). The cheliceral flagellum of pseudoscorpions. Journal of Natural History 1: 393–397. -- Show included taxa

Bologna, M.A. & Taglianti, A.V. (1985). Fauna cavernicola delle Alpi Liguri. Annali del Museo Civico di Storia Naturale di Genova, Supplemento 84: 1–389. -- Show included taxa

Bristowe, W.S. (1929a). The spiders of Cambridgeshire (including harvest spiders and pseudoscorpions). In: Fauna List No. 2 Cambridge, Cambridge Natural History Society: 1–25. -- Show included taxa

Bristowe, W.S. (1938). Arachnida. In: Salzman, L.F. (ed.) The Victoria history of the county of Cambridgeshire and the Isle of Ely Oxford University Press, London 1: 206–212. -- Show included taxa

Callaini, G. (1980a). Considerazioni sugli pseudoscorpioni dell'altopiano del Cansiglio (Notulae Chernetologicae. IV). Animalia, Catania 6: 219–241. -- Show included taxa

Callaini, G. (1983d). Osservazioni sulla fauna Chernetologica di alcuni Rilievi Abruzzesi. Notulae Chernetologicae XIII.. Bollettino del Museo Civico di Storia Naturale, Verona 10: 221–239. -- Show included taxa

Callaini, G. (1986e). Pseudoscorpioni dell'Italia settentrionale nel Museo Civico di Storia Naturale di Verona (Arachnida). Notulae Chernetologicae XIX. Bollettino del Museo Civico di Storia Naturale, Verona 12: 229–255. -- Show included taxa

Canestrini, G. (1875a). Intorno ai Chernetidi ed Opilionidi della Calabria. Atti della Società Veneto-Trentina di Scienze Naturali Residente in Padova 4: 1–12. -- Show included taxa

Canestrini, J. (1885). Chernetides Italici. In: Berlese, A. (ed.) Acari, Myriapoda et Scorpiones hucusque in Italia reperta A. Berlese, Padova fascicolo XIX. -- Show included taxa

Caporiacco, L. di (1926). Secondo saggio sulla fauna aracnologica della Carnia e regioni limitrofe. Memorie della Società Entomologica Italiana 5: 70–130. -- Show included taxa

Caporiacco, L. di (1950a). Gli Aracnidi della laguna di Venezia. II Nota. Bollettino della Società Veneziana di Storia Naturale e del Museo Civico di Storia Naturale 5: 114–140. -- Show included taxa

Caporiacco, L. di (1951b). Aracnidi cavernicoli Liguri. Annali del Museo Civico di Storia Naturale di Genova 64: 101–110. -- Show included taxa

Carpenter, G.H. (1902). Arachnida. In: Bigger, F.J., Praeger, R.L. and Vinycomb, J. (ed.) Guide to Belfast and the counties of Down and Antrim M'Caw, Stevenson and Orr, Belfast: 217–220. -- Show included taxa

Carpenter, G.H. & Evans, W. (1899). Additional records of spiders and other arachnids from the Edinburgh district (second instalment). Proceedings of the Royal Physical Society of Edinburgh 14: 168–181. -- Show included taxa

Cerruti, M. (1959). Aggiunta al i elenco della fauna cavernicola del Lazio e delle regioni limitrofe (Toscana esclusa). Fragmenta Entomologica 3: 49–63. -- Show included taxa

Chamberlin, J.C. (1929a). A synoptic classification of the false scorpions or chela-spinners, with a report on a cosmopolitan collection of the same. Part 1. The Heterosphyronida (Chthoniidae) (Arachnida-Chelonethida). Annals and Magazine of Natural History (10) 4: 50–80. -- Show included taxa

Christophoryová, J., Štáhlavský, F. & Fedor, P. (2011). An updated identification key to the pseudoscorpions (Arachnida: Pseudoscorpiones) of the Czech Republic and Slovakia. Zootaxa 2876: 35–48. -- Show included taxa

Christophoryová, J., Štáhlavský, F., Krumpál, M. & Fedor, P. (2012). Pseudoscorpions of the Czech Republic and Slovakia: an annotated and revised checklist (Arachnida: Pseudoscorpiones). North-Western Journal of Zoology 8(1): 1–21. -- Show included taxa

Cooreman, J. (1946b). Note sur les pseudoscorpions de la faune Belge. Bulletin du Musée Royal d'Histoire Naturelle de Belgique 22(2): 1–8. -- Show included taxa

Costantini, G.P. (1976). Gli scorpioni e pseudoscorpioni della provincia di Brescia. Note di Aracnologia. II. Natura, Bresciana 13: 121–124. -- Show included taxa

Crocker, J. (1978). Introduction to false scorpions. British Arachnological Society, Secretary's News Letter 21: 4–9. -- Show included taxa

Ćurčić, B.P.M. (1974a). Arachnoidea. Pseudoscorpiones. In: Catalogus faunae Jugoslaviae Academie Slovène, Ljubljana 3(4): 1–35. -- Show included taxa

Ćurčić, B.P.M. (1976b). Une contribution a la connaissance de la faune des pseudoscorpions en Serbie. Glasnik Muzeja Srpske Zemlje, Beograd (B) 31: 169–184. -- Show included taxa

Ćurčić, B.P.M., Dimitrijević, R.N. & Legakis, A. (2004). The pseudoscorpions of Serbia, Montenegro, and the Republic of Macedonia. Institute of Zoology, Faculty of Biology, University of Belgrade, Belgrade, Athens. -- Show included taxa

Daday, E. (1889a). A Magyar Nemzeti Muzeum álskorpióinak áttekintése. Természetrajzi Füzetek 11: 111–136, 165–192. -- Show included taxa

Daday, E. (1889d). Ujabb adatok o magyar-fauna álskorpióinak ismeretéhez. Természetrajzi Füzetek 12: 25–28. -- Show included taxa

Daday, E. (1889e). Adatok a Balkán-félsziget álskorpió-faunájának ismeretéhez. Természetrajzi Füzetek 12: 80–84, 113. -- Show included taxa

Daday, E. (1918). Ordo Pseudoscorpiones. In: A Magyar Birodalom Allatvilága Regia Societas Scientiarum Naturalium Hungarica, Budapest: 1–2. -- Show included taxa

Delfosse, E. (2003b). Catalogue préliminaire des Pseudoscorpions de France métropolitaine (Arachnida Pseudoscorpiones). Bulletin de Phyllie 17: 24–48. -- Show included taxa

Drogla, R. & Blick, T. (1996). Rote Liste der Pseudoskorpione Deutschlands (Arachnida: Pseudoscorpiones). Arachnologische Mitteilungen 11: 36–38. -- Show included taxa

Ducháč, V. (1989a). Příspěvek k faunistice štírků Československa (Pseudoscorpionidea). Zbornik Slovenskeho Národneho Múzea Prirodovedny 35: 179–182. -- Show included taxa

Ducháč, V. (1994). Faunisticko-bionomické poznámky k některým druhům štírků České Republiky a Slovenské Republiky. Fauna Bohemiae Septentrionalis 19: 139–153. -- Show included taxa

Ducháč, V. (1999a). The contemporary research of the pseudoscorpion fauna in the Czech Republic. American Arachnology 59: 14. -- Show included taxa

El-Hennawy, H.K. (1988b). Pseudoscorpions of Egypt, key and list of species (Arachnida: Pseudoscorpionida). Serket 1(3): 9–18. -- Show included taxa

Ellingsen, E. (1909a). Contributions to the knowledge of the pseudoscorpions from material belonging to the Museo Civico in Genova. Annali del Museo Civico di Storia Naturale di Genova (3) 4: 205–220. -- Show included taxa

Ellingsen, E. (1910a). Die Pseudoskorpione des Berliner Museums. Mitteilung aus dem Zoologischen Museum in Berlin 4: 357–423. -- Show included taxa

Evans, G.O. & Browning, E. (1954). Synopses of the British Fauna. No. 10. Pseudoscorpiones. Linnean Society, London. -- Show included taxa

Evans, W. (1901a). Roncus cambridgii, L. K., and other chernetids in Scotland.. Annals of Scottish Natural History 10: 53–54. -- Show included taxa

Fowles, A. (1994). A provisional checklist of the invertebrates recorded from Wales. 1. False scorpions, harvestmen and spiders (Arachnida). Cyngor Cefn Gwlad Cymru (Countryside Council for Wales), Bangor, Gwynedd. -- Show included taxa

Franciscolo, M. (1955). Fauna cavernicola del Savonese. Annali del Museo Civico di Storia Naturale di Genova 67: 1–223. -- Show included taxa

Gardini, G. (1975). Pseudoscorpioni dell'Isola di Capraia (Arcipelago Toscano) (Arachnida). Lavori della Societa Italiana di Biogeografia, nuova serie 5: 1–12. -- Show included taxa

Gardini, G. (1980c). Catalogo degli Pseudoscorpioni cavernicoli italiani (Pseudoscorpioni d'Italia VIII). Memorie della Società Entomologica Italiana 58: 95–140. -- Show included taxa

Gardini, G. (1991a). Pseudoscorpioni cavernicoli del Veneto (Arachnida). (Pseudoscorpioni d'Italia XIX). Bollettino del Museo Civico di Storia Naturale, Verona 15: 167–214. -- Show included taxa

Gardini, G. (1995a). Arachnida Pseudoscorpionida. In: Minelli, A, Ruffo, S. and La Posta, S. (ed.) Checklist delle specie della fauna Italiana Calderini, Bologna 22: 1–8. -- Show included taxa

Gardini, G. (2000). Catalogo degli Pseudoscorpioni d'Italia (Arachnida). Fragmenta Entomologica 32, Supplemento: 1–181. -- Show included taxa

Goddard, S.J. (1976b). Population dynamics, distribution patterns and life cycles of Neobisium muscorum and Chthonius orthodactylus (Pseudoscorpiones: Arachnida). Journal of Zoology, London 178: 295–304. -- Show included taxa

Goddard, S.J. (1979a). The population metabolism and life history tactics of Neobisium muscorum (Leach) (Arachnida: Pseudoscorpiones). Oecologia 42: 91–105. -- Show included taxa

Goddard, S.J. (1979b). A modified heat extractor for separating arthropods from deciduous woodland litter. Pedobiologia 19: 26–33. -- Show included taxa

Godfrey, R. (1908). The false scorpions of Scotland. Annals of Scottish Natural History 17: 90–100, 155–161. -- Show included taxa

Godfrey, R. (1910). The false scorpions of Scotland (continued). Annals of Scottish Natural History 19: 23–33. -- Show included taxa

Groh, K. (2007). Afterskorpione - pseudoscorpions - Arachnida, Pseudoscorpiones (Inventaire de la biodiversité dans la forêt "Schnellert" (Commune de Berdorf)). Ferrantia 50: 205–207. -- Show included taxa

Hagen, H. (1870). Synopsis pseudoscorpionidum synonymica. Proceedings of the Boston Society of Natural History 13: 263–272. -- Show included taxa

Harvey, M.S. (1991a). Catalogue of the Pseudoscorpionida. Manchester University Press, Manchester. -- Show included taxa

Harvey, P.R. (1992). Pseudoscorpions. Essex Naturalist 11: 149–150. -- Show included taxa

Henderickx, H. (1999b). Naamlijst van de Belgische pseudoschorpioenen (Arachnida Pseudoscorpionida). Bulletin de la Societe Royale Belge d'Entomologie 135: 66–71. -- Show included taxa

Howes, C.A. (1972a). A review of Yorkshire pseudoscorpions. Naturalist, Hull 918: 107–110. -- Show included taxa

Inzaghi, S. (1981). Pseudoscorpioni raccolti dal Sig. M. Valle in nidi di Talpa europaea L. nella provincia di Bergamo con descrizione di una nuova specie del gen. Chthonius C.L. Koch.. Bollettino della Societa Entomologica Italiana 113: 67–73. -- Show included taxa

Ionescu, M.A. (1936). Contribuţiuni la studiul Pseudoscorpionilor din România. Buletinul Societaţii Naturaliştilor din România 8: 1–5. -- Show included taxa

Isaia, M., Paschetta, M., Lana, E., Pantini, P., Schönhofer, A.L., Christian, E. & Badino, G. (2011). Aracnidi sotterranei delle Alpi Occidentali italiane / Subterranean arachnids of the western Italian Alps (Arachnida: Araneae, Opiliones, Palpigradi, Pseudoscorpiones). Museo Regionale di Scienze Naturali, Torino, Italia Monografie XLVII: i–xi, 1–325. -- Show included taxa

Jackson, A.R. (1910). On some arthropods observed in 1909. Lancashire Naturalist (new series) 3: 17–22, 46–51. -- Show included taxa

Jones, P.E. (1980c). Provisional atlas of the Arachnida of the British Isles. Part 1. Pseudoscorpiones.. Institute of Terrestrial Ecology, Monks Wood. -- Show included taxa

Judson, M.L.I. (1979a). Pseudoscorpions in Hertfordshire. Transactions of the Hertfordshire Natural History Society and Field Club 28: 58–64. -- Show included taxa

Judson, M.L.I. (1987). Further records of pseudoscorpions (Arachnida) from Hertfordshire. Transactions of the Hertfordshire Natural History Society and Field Club 29: 368–370. -- Show included taxa

Kárpáthegyi, P. (2007a). Pseudoscorpions of Hungary. Folia Historico Naturalia Musei Matraensis 31: 81–90. -- Show included taxa

Kárpáthegyi, P. (2007b). Check list of the Hungarian Pseudoscorpiones. Folia Historico Naturalia Musei Matraensis 31: 91–97. -- Show included taxa

Kästner, A. (1928). 2. Ordnung: Moos- oder Afterskorpione, Pseudoscorpiónes Latr. (Chernétes Simon; Chelonéti Thorell; Chernetídea Camb.). In: Brohmer, P, Ehrmann, P. and Ulmer, G. (ed.) Die Tierwelt Mitteleuropas Quelle und Meyer, Leipzig 3 (1, IV): 1–13. -- Show included taxa

Kew, H.W. (1911a). A synopsis of the false scorpions of Britain and Ireland. Proceedings of the Royal Irish Academy, section B 29: 38–64. -- Show included taxa

Kew, H.W. (1916a). An historical account of the pseudoscorpion-fauna of the British islands. Journal of the Quekett Microscopical Club (2) 13: 117–136. -- Show included taxa

Kew, H.W. (1930). On the spermatophores of the Pseudoscorpiones Chthonius and Obisium. Proceedings of the Zoological Society of London 100: 253–256. -- Show included taxa

Koch, C.L. (1843). Die Arachniden. Getreu nach der Natur Abgebildet und Beschrieben. C.H. Zeh'schen, Nürnberg 10. -- Show included taxa

Koch, L. (1873). Uebersichtliche Dartstellung der Europäischen Chernetiden (Pseudoscorpione). Bauer und Raspe, Nürnberg. -- Show included taxa

Koch, L. (1877). Verzeichniss der bei Nürnberg bis jetzt beobachteten Arachniden (mit Ausschluss der Ixodiden und Acariden) und Beschreibungen von neuen, hier vorkommenden Arten. Abhandlungen der Naturhistorischen Gesellschaft zu Nürnberg 6: 113–198. -- Show included taxa

Krajčovičová, K., Christophoryová, J. & Krumpál, M. (2013). Pseudoscorpions (Arachnida: Pseudoscorpiones) of the southwestern part of Český Kras Protected Landscape, Czech Republic. Munis Entomology and Zoology 8: 714–725. -- Show included taxa

Krumpál, M. (1980). Šťúriky (Pseudoscorpionidea) gaderskej doliny (Vel'ká Fatra). Entomologicke Problemy 16: 23–29. -- Show included taxa

Krumpál, M. & Kiefer, M. (1981). Príspevok k Poznaniu Šťúrikov Čel'ade Chthoniidae v ČSSR (Pseudoscorpionidea). Zprávy Československé Společnosti Entomologické Pri CSAV, Praha 17: 127–130. -- Show included taxa

Lazzeroni, G. (1969a). Sur la faune de pseudoscorpions de la région apenninique méridionale. (Recherches sur les Pseudoscorpions. III.). Memorie del Museo Civico di Storia Naturale di Verona 16: 321–344. -- Show included taxa

Lazzeroni, G. (1969b). Contributo alla conoscenza degli pseudoscorpioni della regione Veronese. (Ricerche sugli Pseudoscorpioni. IV). Memorie del Museo Civico di Storia Naturale di Verona 16: 379–418. -- Show included taxa

Lazzeroni, G. (1969c). Ricerche sugli Pseudoscorpioni. VI. Il popolamento della Sardegna. Fragmenta Entomologica 6: 223–251. -- Show included taxa

Lazzeroni, G. (1970a). Ricerche sugli Pseudoscorpioni. III. Considerazioni biogeografiche sulla fauna della regione appenninica meridionale. Bulletin du Muséum National d'Histoire Naturelle, Paris (2) 41(supplément 1): 205–208. -- Show included taxa

Lazzeroni, G. (1970c). Ricerche sugli Pseudoscorpioni. V. L'isola di Giannutri. Memorie, Atti della Societa Toscana di Scienze Naturali (B) 76: 101–112. -- Show included taxa

Leach, W.E. (1817). The zoological miscellany; being descriptions of new or interesting animals. Nodder, London 3. -- Show included taxa

Leclerc, P. & Heurtault, J. (1979). Pseudoscorpions de l'Ardèche. Revue Arachnologique 2: 239–247. -- Show included taxa

Legg, G. (1971a). False-scorpions: the families Chthoniidae, Neobisiidae and Cheiridiidae. Countryside 21: 472–477. -- Show included taxa

Legg, G. (1973b). The structure of encysted sperm of some British Pseudoscorpiones (Arachnida). Journal of Zoology, London 170: 429–440. -- Show included taxa

Legg, G. (1975a). A generalised account of the male genitalia and associated glands of pseudoscorpions (Arachnida). Bulletin of the British Arachnological Society 3: 66–74. -- Show included taxa

Legg, G. (1975b). The genitalia and associated glands of five British species belonging to the family Chthoniidae (Pseudoscorpiones: Arachnida). Journal of Zoology, London 177: 99–121. -- Show included taxa

Legg, G. & Jones, R.E. (1988). Synopses of the British fauna (new series). 40. Pseudoscorpions (Arthropoda; Arachnida). Brill/Backhuys, Leiden. -- Show included taxa

Legg, G. & O'Connor, J.P. (1997). A review of the Irish pseudoscorpions (Arachnida: Pseudoscorpiones). Bulletin of the Irish Biogeographical Society 20: 105–126. -- Show included taxa

Lucas, H. (1840). Histoire naturelle des Crustacés, des Arachnides et des Myriapodes. P. Duménil, Paris. -- Show included taxa

Mahnert, V. (2004). Die Pseudoskorpione Österreichs (Arachnida, Pseudoscorpiones). Denisia 12: 459–471. -- Show included taxa

Mahnert, V. (2011d). Pseudoscorpiones (Arachnida). In: Schuster, R. (ed.) Checklisten der Fauna Österreichs Verlag der Österreichischen Akademie der Wissenschaften, Wien 5: 28–39. -- Show included taxa

Mahnert, V. & Weber, D. (2013). Pseudoskorpione (Arachnida, Pseudoscorpiones) aus Höhlen des Grossherzogtums Luxemburg. Ferrantia 69: 108–114. -- Show included taxa

Marcuzzi, G. (1968). Osservazioni ecologiche sulla fauna del suolo di alcune regioni forestali Italiane. Annali del Centro di Economia Montana delle Venezie 7: 205–331. -- Show included taxa

Matsuda, R. (1979). Abnormal metamorphosis and arthropod evolution. In: Gupta, A.P. (ed.) Arthropod phylogeny Van Nostrand Reinhold, New York: 137–256. -- Show included taxa

McIntire, S.J. (1868). On pseudo-scorpiones. Journal of the Quekett Microscopical Club 1: 8–14. -- Show included taxa

McIntire, S.J. (1869). Pseudoscorpions. Hardwicke's Science Gossip 5: 243–247. -- Show included taxa

Morley, C. (1930). Arachnida. Transactions of the Suffolk Naturalists' Society 1: 140–141. -- Show included taxa

Morley, C. (1940). The remaining Arachnida. Transactions of the Suffolk Naturalists' Society 4: 165–174. -- Show included taxa

Ozimec, R. (2004). List of Croatian pseudoscorpion fauna (Arachnida, Pseudoscorpiones). Natura Croatica 13: 381–394. -- Show included taxa

Palmgren, P. (1973). Über die Biotopverteilung waldbodenlebender Pseudoscorpionidea (Arachnoidea) in Finnland und Österreich. Commentationes Biologicae 61: 1–11. -- Show included taxa

Pavesi, P. (1884). Materiali per lo studio della fauna Tunisina raccolti da G. e L. Doria. Annali del Museo Civico di Storia Naturale di Genova 20: 446–486. -- Show included taxa

Petrov, B.P. & Šťáhlavský, F. (2007). New species of pseudoscorpions (Arachnida: Pseudoscorpiones) for the fauna of Bulgaria. Historia Naturalis Bulgarica 18: 15–27. -- Show included taxa

Pickard-Cambridge, O. (1892). On the British species of false-scorpions. Proceedings of the Dorset Natural History and Antiquarian Field Club and Archaeological Society 13: 199–231. -- Show included taxa

Platen, R., Blick, T., Bliss, P., Drogla, R., Malten, A., Martens, J., Sacher, P. & Wunderlich, J. (1995). Verzeichnis der Spinnentiere (excl. Acarida) Deutschlands (Arachnida: Araneida, Opilionida, Pseudoscorpionida). Arachnologische Mitteilungen, Sonderband 1: 1–55. -- Show included taxa

Roewer, C.F. (1937). Chelonethi oder Pseudoskorpione. In: Bronns, H.G. (ed.) Bronn's Klassen und Ordnungen des Tierreichs Akademische Verlagsgesellschaft M.B.H., Leipzig 5(IV)(6)(1): 161–320. -- Show included taxa

Rundle, A.J. (1979). Pseudoscorpions and harvestmen in Bedfordshire. Bedfordshire Naturalist 33: 47–50. -- Show included taxa

Salmon, S.J. (1972). Recent records of British pseudoscorpions. Bulletin of the British Arachnological Society 2: 66–68. -- Show included taxa

Savory, T.H. (1945). The spiders and allied orders of the British Isles comprising descriptions of every family of British spiders, every species of harvestmen and false scorpions also the more familiar of the British mites and sea-spiders. Frederick Warne & Co., London. [2nd edition] -- Show included taxa

Schawaller, W. (1981b). Pseudoskorpione von Korsika (Arachnida, Pseudoscorpionidea). Entomologica Basiliensia 6: 42–51. -- Show included taxa

Schenkel, E. (1928). Pseudoscorpionida (Afterskorpione). In: Dahl, F. (ed.) Die Tierwelt Deutschlands G. Fischer, Jena 8: 52–72. -- Show included taxa

Simon, E. (1879a). Les Ordres des Chernetes, Scorpiones et Opiliones. In: Les Arachnides de France Librairie Encyclopédique de Roret, Paris 7: 1–332. -- Show included taxa

Simon, E. (1885c). Étude sur les Arachnides recueillis en Tunisie en 1883 et 1884 par Mm. A. Letourneux, M. Sédillot et Valery Mayet. In: Exploration Scientifique de la Tunisie Imprimerie Nationale, Paris: 1–55. -- Show included taxa

Simon, E. (1898d). Studio sui Chernetes Italiani conservati nel Museo Civico di Genova con descrizione di nuova specie. Annali del Museo Civico di Storia Naturale di Genova (2) 19: 20–24. -- Show included taxa

Simon, E. (1900b). Studio sui Chernetes Italiani conservati nel Museo Civico di Genova. II. Annali del Museo Civico di Storia Naturale di Genova (2) 20: 593–595. -- Show included taxa

Šťáhlavský, F. (2001). Štírci (Arachnida: Pseudoscorpiones) Prahy. Klapalekiana 37: 73–121. -- Show included taxa

Šťáhlavský, F. (2011). Štírci (Arachnida: Pseudoscorpiones) CHKO Třeboňsko a okolí. Klapalekiana 47: 247–258. -- Show included taxa

Šťáhlavský, F. & Chytil, J. (2013). Štírci (Arachnida: Pseudoscorpiones) Biosférické rezervace Dolní Morava a okolí (Česká republika). Klapalekiana 49: 73–88. -- Show included taxa

Šťáhlavský, F. & Král, J. (2004). Karyotype analysis and achiasmatic meiosis in pseudoscorpions of the family Chthoniidae (Arachnida: Pseudoscorpiones). Hereditas 140: 49–60. -- Show included taxa

Šťáhlavský, F. & Krásný, L. (2007). Štírci (Arachnida: Pseudoscorpiones) Dolního Povltaví a Podřipska. Bohemia Centralia, Praha 28: 427–436. -- Show included taxa

Szalay, L. (1968). Pókszabásúak I. Arachnoidea I. In: Magyarország Allatvilága (Faunae Hungariae) Akadémiai Kiadó, Budapest 18(1): 1–122. -- Show included taxa

Théis, C. de (1832). Lettre adressée à M. Audouin, sur quelques Arachnides des genres Hydrachna et Chelifer. Annales des Sciences Naturelles 27: 57–78. -- Show included taxa

Tömösváry, Ö. (1883). Pseudoscorpiones faunae Hungaricae. A Magyar fauna álskorpiói. Magyar Tudományos Akadémia Matematikai és Természettudományi Közlemények 18: 135–256. -- Show included taxa

Tooren, D. van den (2005b). Naamlijst en determinatiesleutel tot de Pseudoschorpioenen van Nederland (Arachnida: Pseudoscorpiones). Nederlandse Faunistische Mededelingen 23: 91–102. -- Show included taxa

Tulk, A. (1844). Note upon Obisium orthodactylum (Leach). Annals and Magazine of Natural History (1) 13: 55–57. -- Show included taxa

Vachon, M. (1934g). Pseudoscorpionides de la Côte-d'Or (2e liste). Bulletin Scientifique de Bourgogne 4: 162–163. -- Show included taxa

Verner, P.H. (1971). Štírci - Pseudoscorpionidea. In: Daniel, M. and Černý, V. (ed.) Klič Zviřeny ČSSR Českoslovenká Akademie Věd, Praha 4: 19–31. -- Show included taxa

Warburton, C. (1909). Arachnida Embolobranchiata (scorpions, spiders, mites, etc.). In: Harmer, S.F. and Shipley, A.E. (ed.) The Cambridge natural history MacMillan and Co., London 4: 297–474. -- Show included taxa

Cawley, M. (2016). An indexed bibliographical checklist of the false-scorpions (Arachnida: Pseudoscorpiones) of Ireland (1836-2014). Bulletin of the Irish Biogeographical Society 40: 116–124. -- Show included taxa

Logs
Date Type Reference Old value
2022-06-18 Taxonomic reference update McIntire, 1869
2022-06-18 Taxonomic reference update McIntire, 1868
2022-04-06 New taxonomic reference entry Cawley, 2016
2022-03-20 New taxonomic reference entry Canestrini, 1875
2022-03-18 Species updated Old value
2021-12-22 New taxonomic reference entry Canestrini, 1875