Chthonius tenuis L. Koch, 1873

SPECIES |   ACCEPTED

Original combination: Chthonius tenuis

LSID urn:lsid:wac.nmbe.ch:name:4897772d-0ffb-42c6-982d-0c364ec19319


Taxonomic references

Chthonius tenuis

Koch, 1873: 51–52 original description
Stecker, 1874a: 238–239
Stecker, 1875d: 88–89
Simon, 1879a: 72–73, 312
Pickard-Cambridge, 1885: 103
Pickard-Cambridge, 1892: 211, plate A figs 17–19
Müller and Schenkel, 1895: 822
Simon, 1898c: 3
Simon, 1898d: 22
Simon, 1899e: 73
Ellingsen, 1907a: 172
Simon, 1907: 551
Gozo, 1908: 132
Jackson, 1908: 73
Bignotti, 1909: 67
Jackson, 1909: 438
Kew, 1909b: 259
Stephenson, 1909: 195
Warburton, 1909: 438
Ellingsen, 1910a: 402
Jackson, 1910: 50
Kew, 1911a: 57–58, fig. 21
Lessert, 1911: 39–40, fig. 30
Ellingsen, 1912c: 174
Krausse-Heldrungen, 1912: 65
Bristowe, 1922: 11
Caporiacco, 1923: 132
Caporiacco, 1926: 75
Jeannel, 1926: 155
Schenkel, 1926: 311
Beier, 1928: 312
Kästner, 1928: 9
Redikorzev, 1928: 136
Schenkel, 1928: 71, fig. 33
Beier, 1929d: 156
Beier, 1929e: 454
Berland, 1929: 18
Kew, 1930: 253–256, fig. 1a
Beier, 1930h: 95
Boldori, 1932: 127
Schenkel, 1933: 17
Vachon, 1934g: 162
Caporiacco, 1936b: 328
Roewer, 1937: 239
Wolf, 1938: 608
Savory, 1945: 180, 182
Caporiacco, 1948c: 240
Cârdeiu, 1948: 4 (as Chtonius tonuis [sic])
Caporiacco, 1951a: 62
Caporiacco, 1951b: 103
Baggini, 1961: 24
Aellen and Strinati, 1962: 46
Pedder, 1965: 108, 109, fig. 1
Strinati, 1966: 326
Zangheri, 1966: 530
Bishop, 1967: 393
Marcuzzi, 1968: 244
Szalay, 1968: 35, figs 23a–b
Gabbutt, 1969c: 231
Lazzeroni, 1970a: 207
Gabbutt, 1970c: 11
Helversen and Martens, 1971: 382, fig. 4
Negrea and Negrea, 1971: 88, 119
Verner, 1971: n/a, fig. 5
Legg, 1973b: 430
Legg, 1975a: 66
Legg, 1975b: 100, figs 1b, 3, 7e, 12a–b
Judson, 1979a: 60
Rundle, 1979: 47
Jones, 1980c: n/a, map 5
Jones, 1983: 396
Jones, 1985: 69
Callaini, 1986d: 524–526, figs 1e-f
Judson, 1987: 369
El-Hennawy, 1988b: 9
Callaini, 1989: 137
Hammen, 1989: n/a, fig. 2b
Hansen, 1991: 8
Fowles, 1994: 17
Fowles, 1996: 6
Judson, 1997a: 43
Ducháč, 1999a: 14
Krumpál and Krumpálova, 2003: 119
Gardini, 2011: 825
Legg and Farr-Cox, 2016: n/a, fig. 5
Muster and Blick, 2016: 548, 551
Andrews, 2017a: 2
Ćurčić, Dimitrijević and Ćurčić, 2020: 18
Červená, Gardini, Jablonski and Christophoryová, 2021: 8, fig. 2A, 3B
Muster, Spelda, Rulik, Thormann, von der Mark and Astrin, 2021: fig. 4, 13819

Chthonius rayi L. Koch:

Simon, 1898d: 22–23 (misidentification)

Chthonius (Chthonius) tenuis L. Koch:

Ellingsen, 1905d: 11–12
Beier, 1932a: 51, fig. 57
Beier, 1939e: 305
Cooreman, 1946b: 3
Beier, 1952e: 2
Beier and Franz, 1954: 454
Evans and Browning, 1954: 10, fig. 8
Beier, 1963a: 256
Beier, 1963b: 31, fig. 23
Beier, 1963h: 150
Beier, 1963i: 283
Ressl, 1965: 289
Rafalski, 1967: 6
Lazzeroni, 1969a: 323
Lazzeroni, 1969c: 224–225
Legg, 1971a: 475, fig. 1c
Lagar, 1972a: 17
Lagar, 1972b: 46
Ćurčić, 1974a: 13
Mahnert, 1975c: 186
Costantini, 1976: 122
Vachon, 1976: 243
Hammen, 1977a: 311
Callaini, 1979b: 134
Callaini, 1979c: 342
Crocker, 1978: 8
Leclerc and Heurtault, 1979: 240
Gardini, 1980c: 101, 120, 121, 123, 124
Inzaghi, 1981: 72
Callaini, 1983c: 280–281
Callaini, 1983d: 222
Callaini, 1983e: 215
Bologna and Taglianti, 1985: 225, fig. 8
Callaini, 1986e: 237
Heurtault, 1986c: 21
Hammen, 1986a: 4, figs 1b, 4b–c, 9a–c, 16a–b
Legg and Jones, 1988: 68–70, figs 13a, 13b(a–f)
Hammen, 1989: n/a, figs 115b–c, 118a–c, 119a–b
Harvey, 1991a: 161–162
Gardini and Oggianu, 1992: 26–28, figs 23–26
Gardini, 1995a: 2
Gardini, 1995b: 45
Platen, Blick, Bliss, Drogla, Malten, Martens, Sacher and Wunderlich, 1995: 54
Drogla and Blick, 1996: 37
Judson and Legg, 1996: 205
Ćurčić in Ćurčić, Dimitrijević, Makarov, Lučić and Karamata, 1997: 32–36, figs 45–73
Gardini, Sabella and Saccone, 1997: 221
Petrov, 1997: 262
Gardini, 2000: 31–33
Henderickx, 1999b: 67
Kovoor and Muñoz-Cuevas, 2000: 65
Šťáhlavský, 2001: 80, fig. 2
Delfosse, 2003b: 29
Ćurčić, Dimitrijević and Legakis, 2004: 236–237
Gardini, 2004: 126
Mahnert, 2004: 464
Šťáhlavský and Král, 2004: 51, figs 1d, 1e, 5d
Šťáhlavský, 2006a: 169
Groh, 2007: n/a, fig. 2
Šťáhlavský and Krásný, 2007: 430
Zaragoza, 2007a: 14
Gardini, 2008a: 630
Gardini, 2008b: 5
Gardini, 2009a: 33–37, 43–51, figs 16–26, 34
Gardini, 2009c: 34
Christophoryová, Štáhlavský and Fedor, 2011: 36, fig. 1K
Isaia, Paschetta, Lana, Pantini, Schönhofer, Christian and Badino, 2011: 207, fig. 5.5
Mahnert, 2011d: 34
Christophoryová, Štáhlavský, Krumpál and Fedor, 2012: 5
Beron, 2015: 49
Ćurčić and Dimitrijević, 2016: 443
Kotrbová, Opatova, Gardini and Šťáhlavský, 2016: 328, 335, fig. 4
Legg, 2019: 190
Červená, Krajčovičová and Christophoryová, 2020b: 221
Novák, 2024: 87

Chthonius (Chthonius) aff. tenuis L. Koch:

Mahnert, 1980d: 22
Gardini, 1991a: 172

Chthonius (Chthonius) sp. prope tenuis L. Koch:

Caoduro, Osella and Ruffo, 1994: 34

Not Chthonius tenuis L. Koch:

Simon, 1900b: 595 (misidentification; see Chthonius ischnocheles (Hermann))
Ellingsen, 1909a: 211, 215 (possible misidentification; see Chthonius lanzai Caporiacco)

Chthonius Ephippiochthonius tenuis L. Koch:


Synonyms
Chthonius (Chthonius) tenuichelatus Hadži, 1937 -- synonymised by Beier, 1963b: 31
LSID urn:lsid:wac.nmbe.ch:name:380ac378-26e9-499c-a63f-f46783ce145a
Chthonius (Chthonius) tenuichelatus
Hadži, 1937: 162–14, figs 6a–g (synonymised by Beier, 1963b: 31) original description
Roewer, 1940: 343
Ćurčić, 1974a: 13
Ćurčić, Dimitrijević and Legakis, 2004: 236
Chthonius microphthalmus vallei Caporiacco, 1949 -- synonymised by Gardini and Oggianu, 1992: 26
LSID urn:lsid:wac.nmbe.ch:name:848baa2b-8827-412b-82a3-34a3eab20bdc
Chthonius microphthalmus vallei
Caporiacco in Bianchi, Caporiacco, Massera and Valle, 1949: 498–499, fig. 1 (synonymised by Gardini and Oggiano, 1992: 26). in Bianchi, Caporiacco, Massera and Valle, 1949: 498–499, fig. 1 (synonymised by Gardini and Oggiano, 1992: 26) original description
Chthonius (Chthonius) microphthalmus vallei Caporiacco:
Beier, 1963b: 35–36
Gardini, 1980c: 100, 128
Harvey, 1991a: 156
Distribution
Distribution table
Albania
Algeria
Austria
Belgium
Bulgaria
Czechia
France (mainland, Corse)
Germany
Greece (mainland)
Italy (mainland, Sardinia, Sicily)
Luxembourg
Malta
Montenegro
North Macedonia
Poland
Romania
Serbia
Slovakia
Spain (mainland)
Switzerland
Tunisia
UK
Type locality: Digne (as "Dryne"), Alpes-de-Haut-Provence, Provence-Alpes-Côtes-d'Azur, France.

Coordinates

Locality Lat Lon
1 Digne 44.083333 6.233333
Type repository
  • undefined Lectotype male, paralectotype male (BMNH 1913.9.1.997–8)
Zoobank LSID

urn:lsid:zoobank.org:act:E2DC144A-0806-41BB-B70F-7478D488E3AF

References

Aellen, V. & Strinati, P. (1962). Nouveaux matériaux pour une faune cavernicole de la Suisse. Revue Suisse de Zoologie 69: 25–66. -- Show included taxa

Andrews, L. (2017a). How to find pseudoscorpions. Newsletter of the British Arachnological Society 138: 2–3. -- Show included taxa

Baggini, A. (1961). Sulla distribuzione della fauna cavernicola Italiana nelle categorie biospeleologiche. Rassegna Speleologica Italiana 13: 1–35. -- Show included taxa

Beier, M. (1928). Die Pseudoskorpione des Wiener Naturhistorischen Museums. I. Hemictenodactyli. Annalen des Naturhistorischen Museums in Wien 42: 285–314. -- Show included taxa

Beier, M. (1929d). Alcuni pseudoscorpioni raccolti da C. Menozzi. Bollettino della Societa Entomologica Italiana 61: 154–156. -- Show included taxa

Beier, M. (1929e). Zoologische Forschungsreise nach den Jonischen Inseln und dem Peloponnes. I. und II. Teil. Sitzungsberichte der Akademie der Wissenschaften in Wien 128: 425–456. -- Show included taxa

Beier, M. (1930h). Due nuovi pseudoscorpioni della Tunisia. Bollettino del Laboratorio di Zoologia Generale e Agraria del R. Istituto Superiore Agrario di Portici 24: 95–98. -- Show included taxa

Beier, M. (1932a). Pseudoscorpionidea I. Subord. Chthoniinea et Neobisiinea. Tierreich 57: i–xx, 1–258. -- Show included taxa

Beier, M. (1939e). Die Pseudoscorpione des oberösterreichischen Landesmuseums in Linz. Jahrbuch des Vereines für Landeskunde und Heimatpflege im Gau Oberdonau, Linz 88: 303–312. -- Show included taxa

Beier, M. (1952e). Ordn.: Pseudoscorpionidea, Afterskorpione. In: Strouhal, H. (ed.) Catalogus faunae Austriae Springer-Verlag, Wien 9a: 2–6. -- Show included taxa

Beier, M. (1963a). Sizilianische Pseudoscorpione. Bolletino Accademia Gioenia di Scienze Naturali in Catania (4) 7: 253–263. -- Show included taxa

Beier, M. (1963b). Ordnung Pseudoscorpionidea (Afterskorpione). In: Bestimmungsbücher zur Bodenfauna Europas Akademie-Verlag, Berlin 1: vi, 313 pp.. -- Show included taxa

Beier, M. (1963h). Pseudoscorpione aus dem Museum "Enrico Caffi" in Bergamo. Rendiconti dell'Instituto Lombardo di Scienze e Lettere 97B: 147–156. -- Show included taxa

Beier, M. (1963i). Appenninische Pseudoscorpione. Memorie del Museo Civico di Storia Naturale di Verona 10: 283–286. -- Show included taxa

Beier, M. & Franz, H. (1954). 16. Ordnung: Pseudoscorpionidea. In: Franz, H. (ed.) Die Nordost-Alpen im Spiegel ihrer Landtierwelt Universitätsverlag Wagner, Innsbruck 1: 453–459. -- Show included taxa

Berland, L. (1929). Arachnides et Crustacès. In: Perrier, R. (ed.) Faune de la France en tableaux synoptiques Librairie Delagrave, Paris 2: 1–220. -- Show included taxa

Beron, P. (2015). Cave fauna of Bulgaria. East West, Sofia. -- Show included taxa

Bianchi, C., Caporiacco, L. di, Massera, M.G. & Valle, A. (1949). Raccolte faunistiche della Grotta della Spipola (Bologna). Commentationes Pontificiae Academia Scientarum 13: 493–527. -- Show included taxa

Bignotti, G. (1909). Elenco dei Pseudoscorpioni trovati in Italia e loro distribuzione geografica. Atti della Società dei Naturaliste e Matematici, Modena (4) 11: 56–76. -- Show included taxa

Bishop, C.P. (1967). The cheliceral flagellum of pseudoscorpions. Journal of Natural History 1: 393–397. -- Show included taxa

Boldori, L. (1932). Altri quattro anni di ricerche nelle caverne italiane. Grotte d'Italia 6: 111–129. -- Show included taxa

Bologna, M.A. & Taglianti, A.V. (1985). Fauna cavernicola delle Alpi Liguri. Annali del Museo Civico di Storia Naturale di Genova, Supplemento 84: 1–389. -- Show included taxa

Bristowe, W.S. (1922). Spiders found in the neighbourhood of Oxshott. Proceedings of the South London Entomological and Natural History Society 1922: 1–11. -- Show included taxa

Callaini, G. (1979b). Osservazioni su alcuni pseudoscorpioni delle Isole Eolie (Notulae Chernetologicae II). Redia 62: 129–145. -- Show included taxa

Callaini, G. (1979c). Notulae Chernetologicae. III. Gli pseudoscorpioni della Farma (Arachnida).. Redia 62: 339–354. -- Show included taxa

Callaini, G. (1983c). Notulae Chernetologicae XII. Nuovi reperti sugli pseudoscorpioni della Sardegna. Lavori della Società Italiana di Biogeografia 8: 279–322. -- Show included taxa

Callaini, G. (1983d). Osservazioni sulla fauna Chernetologica di alcuni Rilievi Abruzzesi. Notulae Chernetologicae XIII.. Bollettino del Museo Civico di Storia Naturale, Verona 10: 221–239. -- Show included taxa

Callaini, G. (1983e). Contributo alla conoscenza degli Pseudoscorpioni d'Algeria (Arachnida). Notulae Chernetologicae XVI. Animalia, Catania 10: 211–235. -- Show included taxa

Callaini, G. (1986d). Note sugli pseudoscorpioni raccolti in alcune grotte della Toscana settentrionale (Arachnida) (Notulae Chernetologicae XXV). Redia 69: 523–542. -- Show included taxa

Callaini, G. (1986e). Pseudoscorpioni dell'Italia settentrionale nel Museo Civico di Storia Naturale di Verona (Arachnida). Notulae Chernetologicae XIX. Bollettino del Museo Civico di Storia Naturale, Verona 12: 229–255. -- Show included taxa

Callaini, G. (1989). Il popolamento delle isole Egadi. Un esempio dell'interesse biogeografico degli Pseudoscorpioni (Arachnida). (Notulae Chernetologicae, XXIX). Annali del Museo Civico di Storia Naturale di Genova 87: 137–148. -- Show included taxa

Caoduro, G., Osella, G. & Ruffo, S. (1994). La fauna cavernicola della regione Veronese. Memorie del Museo Civico di Storia Naturale di Verona (IIa serie), Sezione Scienze della Vita (A: Biologica) 11: 1–144. -- Show included taxa

Caporiacco, L. di (1923). Una nuova specie di Chernetide italiano. Bollettino della Societa Entomologica Italiana 55: 131–134. -- Show included taxa

Caporiacco, L. di (1926). Secondo saggio sulla fauna aracnologica della Carnia e regioni limitrofe. Memorie della Società Entomologica Italiana 5: 70–130. -- Show included taxa

Caporiacco, L. di (1936b). Saggio sulla fauna aracnologica del Casentino, Val d'Arno Superiore e Alta Val Tiberina. In: Festschrift zum 60. Geburtstage von Professor Dr. Embrik Strand Privately Published, Riga 1: 326–369. -- Show included taxa

Caporiacco, L. di (1948c). L'aracnofauna della Romagna in base alle raccolte Zangheri. Redia 34: 237–288. -- Show included taxa

Caporiacco, L. di (1951a). Una raccolta di Aracnidi Umbri. Annali del Museo Civico di Storia Naturale di Genova 64: 62–84. -- Show included taxa

Caporiacco, L. di (1951b). Aracnidi cavernicoli Liguri. Annali del Museo Civico di Storia Naturale di Genova 64: 101–110. -- Show included taxa

Cârdeiu, F. (1948). Beiträge zur Fauna der Pseudoscorpioniden aus dem Urwald «Slătioara» Bezirk Cîmpulung (Bucovina). Revista Ştiinţifică "V. Adamachi" 34: 2–4. -- Show included taxa

Christophoryová, J., Štáhlavský, F. & Fedor, P. (2011). An updated identification key to the pseudoscorpions (Arachnida: Pseudoscorpiones) of the Czech Republic and Slovakia. Zootaxa 2876: 35–48. -- Show included taxa

Christophoryová, J., Štáhlavský, F., Krumpál, M. & Fedor, P. (2012). Pseudoscorpions of the Czech Republic and Slovakia: an annotated and revised checklist (Arachnida: Pseudoscorpiones). North-Western Journal of Zoology 8(1): 1–21. -- Show included taxa

Cooreman, J. (1946b). Note sur les pseudoscorpions de la faune Belge. Bulletin du Musée Royal d'Histoire Naturelle de Belgique 22(2): 1–8. -- Show included taxa

Costantini, G.P. (1976). Gli scorpioni e pseudoscorpioni della provincia di Brescia. Note di Aracnologia. II. Natura, Bresciana 13: 121–124. -- Show included taxa

Crocker, J. (1978). Introduction to false scorpions. British Arachnological Society, Secretary's News Letter 21: 4–9. -- Show included taxa

Ćurčić, B.P.M. (1974a). Arachnoidea. Pseudoscorpiones. In: Catalogus faunae Jugoslaviae Academie Slovène, Ljubljana 3(4): 1–35. -- Show included taxa

Ćurčić, B.P.M., Dimitrijević, R.N. & Legakis, A. (2004). The pseudoscorpions of Serbia, Montenegro, and the Republic of Macedonia. Institute of Zoology, Faculty of Biology, University of Belgrade, Belgrade, Athens. -- Show included taxa

Ćurčić, B.P.M., Dimitrijević, R.N., Makarov, S.E., Lučić, L.R. & Karamata, O.S. (1997). New and little-known false scorpions from the Balkan Peninsula, principally from caves, belonging to the families Chthoniidae and Neobisiidae (Arachnida, Pseudoscorpiones). Monographs of the Faculty of Biology, University of Belgrade 2(2): 1–159. -- Show included taxa

Delfosse, E. (2003b). Catalogue préliminaire des Pseudoscorpions de France métropolitaine (Arachnida Pseudoscorpiones). Bulletin de Phyllie 17: 24–48. -- Show included taxa

Drogla, R. & Blick, T. (1996). Rote Liste der Pseudoskorpione Deutschlands (Arachnida: Pseudoscorpiones). Arachnologische Mitteilungen 11: 36–38. -- Show included taxa

Ducháč, V. (1999a). The contemporary research of the pseudoscorpion fauna in the Czech Republic. American Arachnology 59: 14. -- Show included taxa

El-Hennawy, H.K. (1988b). Pseudoscorpions of Egypt, key and list of species (Arachnida: Pseudoscorpionida). Serket 1(3): 9–18. -- Show included taxa

Ellingsen, E. (1905d). Pseudoscorpions from Italy and southern France conserved in the R. Museo Zoologico in Torino. Bollettino dei Musei di Zoologia e di Anatomia Comparata della R. Università di Torino 20(503): 1–13. -- Show included taxa

Ellingsen, E. (1907a). Notes on pseudoscorpions, British and foreign. Journal of the Quekett Microscopical Club (2) 10: 155–172. -- Show included taxa

Ellingsen, E. (1909a). Contributions to the knowledge of the pseudoscorpions from material belonging to the Museo Civico in Genova. Annali del Museo Civico di Storia Naturale di Genova (3) 4: 205–220. -- Show included taxa

Ellingsen, E. (1910a). Die Pseudoskorpione des Berliner Museums. Mitteilung aus dem Zoologischen Museum in Berlin 4: 357–423. -- Show included taxa

Ellingsen, E. (1912c). Pseudoscorpiones (troisième série). Archives de Zoologie Expérimentale et Générale (5) 10: 163–175. -- Show included taxa

Evans, G.O. & Browning, E. (1954). Synopses of the British Fauna. No. 10. Pseudoscorpiones. Linnean Society, London. -- Show included taxa

Fowles, A. (1994). A provisional checklist of the invertebrates recorded from Wales. 1. False scorpions, harvestmen and spiders (Arachnida). Cyngor Cefn Gwlad Cymru (Countryside Council for Wales), Bangor, Gwynedd. -- Show included taxa

Fowles, A. (1996). Aspects of arachnid recording in Wales. Newsletter of the British Arachnological Society 77: 6–7. -- Show included taxa

Gabbutt, P.D. (1969c). Life-histories of some British pseudoscorpions inhabiting leaf litter. In: Sheals, J.G. (ed.) The soil ecosystem Systematics Association Publication no. 8, London: 229–235. -- Show included taxa

Gabbutt, P.D. (1970c). Sampling problems and the validity of life history analyses of pseudoscorpions. Journal of Natural History 4: 1–15. -- Show included taxa

Gardini, G. (1980c). Catalogo degli Pseudoscorpioni cavernicoli italiani (Pseudoscorpioni d'Italia VIII). Memorie della Società Entomologica Italiana 58: 95–140. -- Show included taxa

Gardini, G. (1991a). Pseudoscorpioni cavernicoli del Veneto (Arachnida). (Pseudoscorpioni d'Italia XIX). Bollettino del Museo Civico di Storia Naturale, Verona 15: 167–214. -- Show included taxa

Gardini, G. (1995a). Arachnida Pseudoscorpionida. In: Minelli, A, Ruffo, S. and La Posta, S. (ed.) Checklist delle specie della fauna Italiana Calderini, Bologna 22: 1–8. -- Show included taxa

Gardini, G. (1995b). Pseudoscorpionida. In: Arthropoda di Lampedusa, Linosa e Pantelleria (Canale di Sicilia, Mar Mediterraneo).. Naturalista Sicilano, supplemento 19: 43–49. -- Show included taxa

Gardini, G. (2000). Catalogo degli Pseudoscorpioni d'Italia (Arachnida). Fragmenta Entomologica 32, Supplemento: 1–181. -- Show included taxa

Gardini, G. (2004). Note faunistiche e tassonomiche sugli pseudoscorpioni (Arachnida) epigei (Pseudoscorpioni d'Italia XXXV). In: Il Monte Pastello. Memorie del Museo Civico di Storia Naturale di Verona, 2. serie, Monografie Naturalistiche 1: 123–133. -- Show included taxa

Gardini, G. (2008a). Ricerche zoologiche della nave oceanografica “Minerva” (C.N.R.) sulle isole circumsarde. XXXI.. Annali del Museo Civico di Storia Naturale "G. Doria", Genova 99: 629–651. -- Show included taxa

Gardini, G. (2008b). Pseudoscorpioni d’Italia XLII. Chthoniidae cavernicoli di Sardegna (Pseudoscorpiones). Memorie della Società Entomologica Italiana 87: 3–31. -- Show included taxa

Gardini, G. (2009a). Chthonius (C.) delmastroi n. sp. delle Alpi occidentali e del Piemonte e ridescrizione di Chthonius (C.) tenuis L. Koch, 1873 e di C. (C.) submontanus Beier, 1963 (Pseudoscorpiones Chthoniidae). Rivista Piemontese di Storia Naturale 30: 25–51. -- Show included taxa

Gardini, G. (2009c). Gli Pseudoscorpioni della Riserva Naturale "Agoraie di Sopra e Moggetto" (Liguria, Genova) (Pseudoscorpiones). Bollettino del'Associazione Romana di Entomologia 64: 33–37. -- Show included taxa

Gardini, G. (2011). Pseudoscorpiones. Conservazione Habitat Invertebrati 5: 824–827. -- Show included taxa

Gardini, G. & Oggianu, A. (1992). Note tassonomiche e faunistiche su alcuni Chthonius della fauna Italiana (Pseudoscorpionida, Chthoniidae). Fragmenta Entomologica 24: 13–34. -- Show included taxa

Gardini, G., Sabella, G. & Saccone, L. (1997). Studi sulle comunita' di Pseudoscorpioni dei sistemi forestali dei Nebrodi (Sicilia nordorientale) (Pseudoscorpionida). Fragmenta Entomologica 29: 213–237. -- Show included taxa

Gozo, A. (1908). Gli aracnidi di caverne italiane. Bollettino della Societa Entomologica Italiana 38: 109–139. -- Show included taxa

Groh, K. (2007). Afterskorpione - pseudoscorpions - Arachnida, Pseudoscorpiones (Inventaire de la biodiversité dans la forêt "Schnellert" (Commune de Berdorf)). Ferrantia 50: 205–207. -- Show included taxa

Hadži, J. (1937). Pseudoskorpioniden aus Südserbien. Glasnik Skopskog Naucnog Drustva 17: 151–187. -- Show included taxa

Hammen, L. van der (1977a). A new classification of Chelicerata. Zoölogische Mededelingen 51: 307–319. -- Show included taxa

Hammen, L. van der (1986a). Comparative studies in Chelicerata IV. Apatellata, Arachnida, Scorpionida, Xiphosura. Zoologische Verhandelingen 226: 1–52. -- Show included taxa

Hammen, L. van der (1989). An introduction to comparative arachnology. SPB Academic Publishing, The Hague. -- Show included taxa

Hansen, H. (1991). Ricerche faunistiche del Museo Civico di Storia Naturale di Venezia nell'isola di Pantelleria. XI - Arachnida: Scorpiones, Pseudoscorpiones, Araneae. Bollettino del Museo Civico di Storia Naturale di Venezia 40: 7–19. -- Show included taxa

Harvey, M.S. (1991a). Catalogue of the Pseudoscorpionida. Manchester University Press, Manchester. -- Show included taxa

Helversen, O. von & Martens, J. (1971). Pseudoskorpione und Weberknechte. Die Wutach 1: 377–385. -- Show included taxa

Henderickx, H. (1999b). Naamlijst van de Belgische pseudoschorpioenen (Arachnida Pseudoscorpionida). Bulletin de la Societe Royale Belge d'Entomologie 135: 66–71. -- Show included taxa

Heurtault, J. (1986c). Pseudoscorpions cavernicoles de France: revue synoptique. Mémoires de Biospéologie 12: 19–32. -- Show included taxa

Inzaghi, S. (1981). Pseudoscorpioni raccolti dal Sig. M. Valle in nidi di Talpa europaea L. nella provincia di Bergamo con descrizione di una nuova specie del gen. Chthonius C.L. Koch.. Bollettino della Societa Entomologica Italiana 113: 67–73. -- Show included taxa

Isaia, M., Paschetta, M., Lana, E., Pantini, P., Schönhofer, A.L., Christian, E. & Badino, G. (2011). Aracnidi sotterranei delle Alpi Occidentali italiane / Subterranean arachnids of the western Italian Alps (Arachnida: Araneae, Opiliones, Palpigradi, Pseudoscorpiones). Museo Regionale di Scienze Naturali, Torino, Italia Monografie XLVII: i–xi, 1–325. -- Show included taxa

Jackson, A.R. (1908). On some rare arachnids captured during 1907. Transactions of the Natural History Society of Northumberland, Durham, and Newcastle-upon-Tyne, new series 3: 49–78. -- Show included taxa

Jackson, A.R. (1909). On some rare arachnids obtained during 1908. Transactions of the Natural History Society of Northumberland, Durham, and Newcastle-upon-Tyne, new series 3: 418–439. -- Show included taxa

Jackson, A.R. (1910). On some arthropods observed in 1909. Lancashire Naturalist (new series) 3: 17–22, 46–51. -- Show included taxa

Jeannel, R. (1926). Faune cavernicole de la France avec une étude des conditions d'existence dans le domaine souterrain. In: Encyclopédique entomologique P. Lechevalier, Paris 7: 1–335. -- Show included taxa

Jones, P.E. (1980c). Provisional atlas of the Arachnida of the British Isles. Part 1. Pseudoscorpiones.. Institute of Terrestrial Ecology, Monks Wood. -- Show included taxa

Jones, R.E. (1983). Some recent Isle of Wight pseudoscorpion records. Proceedings of the Isle of Wight Natural History Archaeological Society 7: 396. -- Show included taxa

Jones, R.E. (1985). The false-scorpions of Norfolk. Transactions of the Norfolk and Norwich Naturalists' Society 27: 67–71. -- Show included taxa

Judson, M.L.I. (1979a). Pseudoscorpions in Hertfordshire. Transactions of the Hertfordshire Natural History Society and Field Club 28: 58–64. -- Show included taxa

Judson, M.L.I. (1987). Further records of pseudoscorpions (Arachnida) from Hertfordshire. Transactions of the Hertfordshire Natural History Society and Field Club 29: 368–370. -- Show included taxa

Judson, M.L.I. (1997a). Catalogue of the pseudoscorpion types (Arachnida: Chelonethi) in the Natural History Museum, London. Occasional Papers on Systematic Entomology 11: 1–54. -- Show included taxa

Judson, M.L.I. & Legg, G. (1996). Discovery of the pseudoscorpion Larca lata (Garypoidea, Larcidae) in Britain. Bulletin of the British Arachnological Society 10: 205–210. -- Show included taxa

Kästner, A. (1928). 2. Ordnung: Moos- oder Afterskorpione, Pseudoscorpiónes Latr. (Chernétes Simon; Chelonéti Thorell; Chernetídea Camb.). In: Brohmer, P, Ehrmann, P. and Ulmer, G. (ed.) Die Tierwelt Mitteleuropas Quelle und Meyer, Leipzig 3 (1, IV): 1–13. -- Show included taxa

Kew, H.W. (1909b). Pseudoscorpiones (false-scorpions). In: Grinling, C. et al. (ed.) Survey and record of Woolwich and West Kent South Eastern Union of Scientific Societies, Woolwich: 258–259. -- Show included taxa

Kew, H.W. (1911a). A synopsis of the false scorpions of Britain and Ireland. Proceedings of the Royal Irish Academy, section B 29: 38–64. -- Show included taxa

Kew, H.W. (1930). On the spermatophores of the Pseudoscorpiones Chthonius and Obisium. Proceedings of the Zoological Society of London 100: 253–256. -- Show included taxa

Koch, L. (1873). Uebersichtliche Dartstellung der Europäischen Chernetiden (Pseudoscorpione). Bauer und Raspe, Nürnberg. -- Show included taxa

Kotrbová, J., Opatova, V., Gardini, G. & Šťáhlavský, F. (2016). Karyotype diversity of pseudoscorpions of the genus Chthonius (Pseudoscorpiones, Chthoniidae) in the Alps. Comparative Cytogenetics 10: 325–345. -- Show included taxa

Kovoor, J. & Muñoz-Cuevas, A. (2000). Diversité des Arachnides dans les îles d'Hyères (Porquerolles et Port-Cros, Var, France). Modifications au cors du XXe siècle. Zoosystema 22: 33–69. -- Show included taxa

Krausse-Heldrungen, A.H. (1912). Sardische Chernetiden. Archiv für Naturgeschichte 21: 65–66. -- Show included taxa

Krumpál, M. & Krumpálova, Z. (2003). Šťúriky - Pseudoscorpiones. In: Mašán, P. and Svatoň, J. (ed.) Pavúkovce Národného Parku Poloniny (Arachnida: Araneae, Pseudoscorpiones, Opiliones, Acari - Parasitiformes) Balada Press, Humenné: 115–126. -- Show included taxa

Lagar, A. (1972a). Contribución al conocimiento de los Pseudoescorpiones de España. I. Miscelanea Zoologica 3: 17–21. -- Show included taxa

Lagar, A. (1972b). Contribución al conocimiento de los Pseudoescorpiones de España. II. Speleon 19: 45–52. -- Show included taxa

Lazzeroni, G. (1969a). Sur la faune de pseudoscorpions de la région apenninique méridionale. (Recherches sur les Pseudoscorpions. III.). Memorie del Museo Civico di Storia Naturale di Verona 16: 321–344. -- Show included taxa

Lazzeroni, G. (1969c). Ricerche sugli Pseudoscorpioni. VI. Il popolamento della Sardegna. Fragmenta Entomologica 6: 223–251. -- Show included taxa

Lazzeroni, G. (1970a). Ricerche sugli Pseudoscorpioni. III. Considerazioni biogeografiche sulla fauna della regione appenninica meridionale. Bulletin du Muséum National d'Histoire Naturelle, Paris (2) 41(supplément 1): 205–208. -- Show included taxa

Leclerc, P. & Heurtault, J. (1979). Pseudoscorpions de l'Ardèche. Revue Arachnologique 2: 239–247. -- Show included taxa

Legg, G. (1971a). False-scorpions: the families Chthoniidae, Neobisiidae and Cheiridiidae. Countryside 21: 472–477. -- Show included taxa

Legg, G. (1973b). The structure of encysted sperm of some British Pseudoscorpiones (Arachnida). Journal of Zoology, London 170: 429–440. -- Show included taxa

Legg, G. (1975a). A generalised account of the male genitalia and associated glands of pseudoscorpions (Arachnida). Bulletin of the British Arachnological Society 3: 66–74. -- Show included taxa

Legg, G. (1975b). The genitalia and associated glands of five British species belonging to the family Chthoniidae (Pseudoscorpiones: Arachnida). Journal of Zoology, London 177: 99–121. -- Show included taxa

Legg, G. & Jones, R.E. (1988). Synopses of the British fauna (new series). 40. Pseudoscorpions (Arthropoda; Arachnida). Brill/Backhuys, Leiden. -- Show included taxa

Lessert, R. de (1911). Pseudoscorpions. Catalogue des Invertébrés de la Suisse 5: 1–50. -- Show included taxa

Mahnert, V. (1975c). Pseudoscorpione von den maltesischen Inseln. Fragmenta Entomologica 11: 185–197. -- Show included taxa

Mahnert, V. (1980d). Pseudoskorpione (Arachnida) aus Höhlen Italiens, mit Bemerkungen zur Gattung Pseudoblothrus. Grotte d'Italia (4a) 8: 21–38. -- Show included taxa

Mahnert, V. (2004). Die Pseudoskorpione Österreichs (Arachnida, Pseudoscorpiones). Denisia 12: 459–471. -- Show included taxa

Mahnert, V. (2011d). Pseudoscorpiones (Arachnida). In: Schuster, R. (ed.) Checklisten der Fauna Österreichs Verlag der Österreichischen Akademie der Wissenschaften, Wien 5: 28–39. -- Show included taxa

Marcuzzi, G. (1968). Osservazioni ecologiche sulla fauna del suolo di alcune regioni forestali Italiane. Annali del Centro di Economia Montana delle Venezie 7: 205–331. -- Show included taxa

Müller, F. & Schenkel, E. (1895). Verzeichnis der Spinnen von Basel und Umgegend. Verhandlungen der Naturforschenden Gesellschaft in Basel 10: 691–824. -- Show included taxa

Negrea, A. & Negrea, Ş. (1971). Sur la synusie du guano des grottes du Banat (Roumanie). Travaux de l'Institute de Spéologie "Émile Racovitza" 10: 81–122. -- Show included taxa

Pedder, I.J. (1965). Abnormal segmentation of the abdomen in six species of British pseudoscorpions. Entomologist 98: 108–112. -- Show included taxa

Petrov, B. (1997). A review of Bulgarian pseudoscorpions (Arachnida, Pseudoscorpionida). In: Zabka, M. (ed.) Proceedings of the 16th European Colloquium of Arachnology Wydawnictwo Wyzszej Skoly Rolniczo-Pedagogicznej, Siedlce: 261–269. -- Show included taxa

Pickard-Cambridge, O. (1885). Pseudoscorpions new to Britain. Naturalist, London 10: 103. -- Show included taxa

Pickard-Cambridge, O. (1892). On the British species of false-scorpions. Proceedings of the Dorset Natural History and Antiquarian Field Club and Archaeological Society 13: 199–231. -- Show included taxa

Platen, R., Blick, T., Bliss, P., Drogla, R., Malten, A., Martens, J., Sacher, P. & Wunderlich, J. (1995). Verzeichnis der Spinnentiere (excl. Acarida) Deutschlands (Arachnida: Araneida, Opilionida, Pseudoscorpionida). Arachnologische Mitteilungen, Sonderband 1: 1–55. -- Show included taxa

Rafalski, J. (1967). Zaleszczotki. Pseudoscorpionidea. In: Katalog Fauny Polski Polska Akademia Nauk, Warszawa 32 (1): 1–34. -- Show included taxa

Redikorzev, V. (1928). Beiträge zur Kenntnis der Pseudoscorpionenfauna Bulgariens. Mitteilungen aus dem Königl. Naturwissenschaftlichen Institut in Sofia 1: 118–141. -- Show included taxa

Ressl, F. (1965). Über Verbreitung, Variabilität und Lebensweise einiger österreichischer Afterskorpione. Deutsche Entomologische Zeitschrift 12: 289–295. -- Show included taxa

Roewer, C.F. (1937). Chelonethi oder Pseudoskorpione. In: Bronns, H.G. (ed.) Bronn's Klassen und Ordnungen des Tierreichs Akademische Verlagsgesellschaft M.B.H., Leipzig 5(IV)(6)(1): 161–320. -- Show included taxa

Roewer, C.F. (1940). Chelonethi oder Pseudoskorpione. In: Bronns, H.G. (ed.) Bronn's Klassen und Ordnungen des Tierreichs Akademische Verlagsgesellschaft M.B.H., Leipzig 5(IV)(6)(1): 321–354. -- Show included taxa

Rundle, A.J. (1979). Pseudoscorpions and harvestmen in Bedfordshire. Bedfordshire Naturalist 33: 47–50. -- Show included taxa

Savory, T.H. (1945). The spiders and allied orders of the British Isles comprising descriptions of every family of British spiders, every species of harvestmen and false scorpions also the more familiar of the British mites and sea-spiders. Frederick Warne & Co., London. [2nd edition] -- Show included taxa

Schenkel, E. (1926). Beitrag zur Kenntnis der Schweizerischen Spinnenfauna. II. Teil. Revue Suisse de Zoologie 33: 301–316. -- Show included taxa

Schenkel, E. (1928). Pseudoscorpionida (Afterskorpione). In: Dahl, F. (ed.) Die Tierwelt Deutschlands G. Fischer, Jena 8: 52–72. -- Show included taxa

Schenkel, E. (1933). Beitrag zur Kenntnis der Schweizerischen Spinnenfauna. V. Teil. Spinnen aus dem Saas-Tal (Wallis) und von der Gegend zwischen Trins und Flims (Graubünden). Revue Suisse de Zoologie 40: 11–29. -- Show included taxa

Simon, E. (1879a). Les Ordres des Chernetes, Scorpiones et Opiliones. In: Les Arachnides de France Librairie Encyclopédique de Roret, Paris 7: 1–332. -- Show included taxa

Simon, E. (1898c). Étude sur les Arachnides de la région des Maures (Var). Feuille des Jeunes Naturalistes (3) 29: 2–4. -- Show included taxa

Simon, E. (1898d). Studio sui Chernetes Italiani conservati nel Museo Civico di Genova con descrizione di nuova specie. Annali del Museo Civico di Storia Naturale di Genova (2) 19: 20–24. -- Show included taxa

Simon, E. (1899e). Liste des Arachnides observés à Lyons-la-Forêt (Eure), en Octobre 1898. Feuille des Jeunes Naturalistes (3) 29(340): 72–73. -- Show included taxa

Simon, E. (1900b). Studio sui Chernetes Italiani conservati nel Museo Civico di Genova. II. Annali del Museo Civico di Storia Naturale di Genova (2) 20: 593–595. -- Show included taxa

Simon, E. (1907). Araneae, Chernetes et Opiliones (première série). Archives de Zoologie Expérimentale et Générale (4) 6: 537–553. -- Show included taxa

Šťáhlavský, F. (2001). Štírci (Arachnida: Pseudoscorpiones) Prahy. Klapalekiana 37: 73–121. -- Show included taxa

Šťáhlavský, F. (2006a). Štírci (Arachnida: Pseudoscorpiones) Národního parku Podyjí. Klapalekiana 42: 167–178. -- Show included taxa

Šťáhlavský, F. & Král, J. (2004). Karyotype analysis and achiasmatic meiosis in pseudoscorpions of the family Chthoniidae (Arachnida: Pseudoscorpiones). Hereditas 140: 49–60. -- Show included taxa

Šťáhlavský, F. & Krásný, L. (2007). Štírci (Arachnida: Pseudoscorpiones) Dolního Povltaví a Podřipska. Bohemia Centralia, Praha 28: 427–436. -- Show included taxa

Stecker, A. (1874a). Zur Kenntnis der Chernetidenfauna Böhmens. Sitzungsberichte der Königliche Böhmischen Gesellschaft der Wissenschaften 8: 227–241. -- Show included taxa

Stecker, A. (1875d). Die Chernetidenfauna Böhmens. Zeitschrift für die Gesammten Naturwissenschaften 11: 87–89. -- Show included taxa

Stephenson, H.B. (1909). Occurrence of Obisium muscorum and Chthonius tenuis near Midhurst. Hastings and East Sussex Naturalist 1: 195. -- Show included taxa

Strinati, P. (1966). Faune cavernicole de la Suisse. Éditions du Centre National de la Recherche Scientifique, Paris. -- Show included taxa

Szalay, L. (1968). Pókszabásúak I. Arachnoidea I. In: Magyarország Allatvilága (Faunae Hungariae) Akadémiai Kiadó, Budapest 18(1): 1–122. -- Show included taxa

Vachon, M. (1934g). Pseudoscorpionides de la Côte-d'Or (2e liste). Bulletin Scientifique de Bourgogne 4: 162–163. -- Show included taxa

Vachon, M. (1976). Quelques remarques sur la Pseudoscorpions (Arachnides) cavernicoles de la Suisse à propos de la description de deux espèces nouvelles: Neobisium (N.) aelleni et Neobisium (N.) strausaki. Revue Suisse de Zoologie 83: 243–253. -- Show included taxa

Verner, P.H. (1971). Štírci - Pseudoscorpionidea. In: Daniel, M. and Černý, V. (ed.) Klič Zviřeny ČSSR Českoslovenká Akademie Věd, Praha 4: 19–31. -- Show included taxa

Warburton, C. (1909). Arachnida Embolobranchiata (scorpions, spiders, mites, etc.). In: Harmer, S.F. and Shipley, A.E. (ed.) The Cambridge natural history MacMillan and Co., London 4: 297–474. -- Show included taxa

Wolf, B. (1938). Animalium cavernarum catalogus. Junk, Gravenhage 3. -- Show included taxa

Zangheri, P. (1966). Repertorio sistematico e topografico della flora e fauna vivente e fossile della Romagna. Museo Civico di Storia Naturale di Verona, Memorie Fuori Serie 1 2: 485–854. -- Show included taxa

Zaragoza, J.A. (2007a). Catálogo de los Pseudoescorpiones de la Península Ibérica e Islas Baleares (Arachnida: Pseudoscorpiones). Revista Ibérica de Aracnología 13: 3–91. -- Show included taxa

Legg, G. (2019). Changes in the British pseudoscorpion fauna over the last 50 years. Arachnology 18: 189–195. -- Show included taxa

Červená, M., Gardini, G., Jablonski, D. & Christophoryová, J. (2021). Checklist of pseudoscorpions (Arachnida, Pseudoscorpiones) of Albania. Zoological Studies 60(17): 1–25. -- Show included taxa

Muster, C., Spelda, J., Rulik, B., Thormann, J., von der Mark, L> & Astrin, J.J. (2021). The dark side of pseudoscorpion diversity: The German Barcode of Life campaign reveals high levels of undocumented diversity in European false scorpions. Ecology and Evolution 11(20): 13815–13829. -- Show included taxa

Legg, G. & Farr-Cox, F. (2016). Illustrated key to the British false scorpions (pseudoscorpions). Field Studies Council: Telford. -- Show included taxa

Ćurčić, N.B. & Dimitrijević, R.N. (2016). Checklist of the pseudoscorpions (Arachnida: Pseudoscorpiones) of Montenegro. Ecologica Montenegrina 7: 439–450. -- Show included taxa

Ćurčić, N.B., Ćurčić, N.B. & Ćurčić, S.B. (2020). Checklist of the pseudoscorpions (Arachnida: Pseudoscorpiones) of Serbia. Arthropoda Selecta 29: 13–27. -- Show included taxa

Muster, C. & Blick, T. (2016). Rote Liste und Gesamtartenliste der Pseudoskorpione (Arachnida: Pseudoscorpiones) Deutschlands.. Naturschutz und Biologische Vielfalt 70: 539–561. -- Show included taxa

Novák, J. (2024). An updated checklist for the pseudoscorpion fauna of Hungary (Arachnida: Pseudoscorpiones). Zootaxa 5433: 51–95. -- Show included taxa

Červená, M., Krajčovičová, K. & Christophoryová, J. (2020b). Updated checklist of pseudoscorpions (Arachnida: Pseudoscorpiones) of Central Europe. Arthropoda Selecta 29: 219–228.. Arthropoda Selecta 29: 219–228. -- Show included taxa

Logs
Date Type Reference Old value
2024-12-18 New taxonomic reference entry Novák, 2024
2024-12-18 New taxonomic reference entry Červená, Krajčovičová and Christophoryová, 2020
2023-04-21 New taxonomic reference entry Muster and Blick, 2016
2022-12-17 New taxonomic reference entry Andrews, 2017
2022-11-18 New taxonomic reference entry Ćurčić, Dimitrijević and Ćurčić, 2020
2022-11-13 New taxonomic reference entry Ćurčić and Dimitrijević, 2016
2022-10-03 New taxonomic reference entry Legg and Farr-Cox, 2016
2022-09-25 Species updated Old value
2022-09-25 New taxonomic reference entry Muster, Spelda, Rulik, Thormann, von der Mark and Astrin, 2021
2022-09-25 New taxonomic reference entry Červená, Gardini, Jablonski and Christophoryová, 2021
2022-09-25 New taxonomic reference entry Legg, 2019
2022-06-18 Taxonomic reference update Ellingsen, 1909
2022-06-18 Taxonomic reference update Simon, 1900
2022-03-18 Species updated Old value