Chthonius ischnocheles (Hermann, 1804)

SPECIES |   ACCEPTED

Original combination: Chelifer ischnochelus

LSID urn:lsid:wac.nmbe.ch:name:5b13db56-9c37-4e2a-b203-bff029f3a703


Taxonomic references

Chelifer ischnochelus

Hermann, 1804: 118–119, plate 5 fig. p, plate 6 fig. 14 original description
Latreille, 1818b: 12, 13, plates 348, 349 (as Chelifer ischnochelos)
Gervais, 1844: 81 (as Chelifer ischnocheles)

Obisium ischnocheles (Hermann):

Lucas, 1840: 449
Hagen, 1870: 270

Obisium cavernarum L. Koch:

Gestro, 1887: 497 (misidentification)

Chthonius microphthalmus Simon:

Simon, 1896: 375 (misidentification)
Simon, 1898d: 23 (misidentification)

Chthonius tenuis L. Koch:

Simon, 1900b: 595 (misidentification)

Chthonius ischnocheles (Hermann):

Chamberlin, 1929a: 71–72, figs 2a, 2g, 2k, 2m-n, 3a, 3d
Chamberlin, 1931a: n/a, figs 4a, 11a, 11t, 15a, 28d, 33o, 34d, 40a–c, 42a, 43a–b, 46f, 52a
Beier, 1932g: n/a, fig. 167
Fage, 1933: 54 (as Chthoniuss [sic] ischnochele [sic])
Vachon, 1934g: 162
Vachon, 1934i: 408
Chamberlin, 1935b: 479
Savory, 1935: n/a, fig. 63ii (as Chthonius ischnochelus [sic])
Vachon, 1935j: 187–188
Roewer, 1936: n/a, figs 16, 70a
Vachon, 1936a: 78
Roewer, 1937: 239
Vachon, 1937f: 128
Schenkel, 1938: 37–38
Vachon, 1938a: 73–79, figs 41–45, 60a–b, 61a
Wolf, 1938: 607
Beier, 1939a: n/a, fig. 166a
Lohmander, 1939b: 285–286, fig. 2
Caporiacco, 1940a: 6
Bristowe, 1941: 327
Pearse, 1943: 464 (misidentification; see Chthonius paludis (Chamberlin))
Savory, 1945: 180, 181–182 (as Chthonius ischnochelus [sic])
Bristowe, 1946: 65
Leleup, 1947: 322
Beier, 1948a: 188
Beier, 1948b: 456
Caporiacco, 1949c: 133
Vachon, 1949: n/a, figs 206c, 207c
Gilbert, 1951a: 548–551, figs 1–2 (as Chthonius ischnochelus [sic])
Balazuc, Miré, Sigwalt and Théodoridès, 1951: 187, 215, 216, 219
Parenzan, 1953: 90 (not seen)
Parenzan, 1954: 99 (not seen)
Betts, 1955: 298
Betts, 1955: 298 (as Chthonius ischnochelus [sic])
George, 1955: 123
Cloudsley-Thompson, 1956a: 71 (as Chthonius ischnochelus [sic])
George, 1957: 80
Cloudsley-Thompson, 1958b: 96
Cloudsley-Thompson, 1959: 179 (as Cthonius [sic] ischnocheles)
George, 1961: 38
Pontin, 1961: 135
Vachon, 1961: 102
Meinertz, 1962: 42–44, figs 5b, 25a–b, 26
Gabbutt and Vachon, 1963: 76–97, figs 1–44
Meinertz, 1964: 388–389, map 1
Savory, 1964a: n/a, fig. 33b
Helversen, 1966a: 133–134
Helversen, 1966b: n/a, figs 1c, 4h, 6a
Weygoldt, 1966a: 48, fig. 6
Weygoldt, 1966d: n/a, fig. 61
Zangheri, 1966: 529–530
Bishop, 1967: 393, figs 1–2
Gabbutt, 1967: 470–478, figs 1–2
Turk, 1967: 153
Cloudsley-Thompson, 1968: 126, 134, 135, 139
Helversen, 1968: n/a, fig. 5
Marcuzzi, 1968: 245, 267, 288
Szalay, 1968: 34–35, fig. 22b–c
Weygoldt, 1968: 111–153, figs 2a–b, 5, 16, 18, 20
Gabbutt, 1969c: 230
Weygoldt, 1969a: 70, 122, 123, figs 112a–d
Gabbutt, 1970c: 7–8, fig. 2
Jones, 1970a: 77–79, figs 1–5
Legg, 1970b: n/a, figs 3(2), 3(7)
Marcuzzi, Lorenzoni and di Castri, 1970: 429–430
Hunt and Legg, 1971: 476–478
Marcuzzi, Dalla Venezia and Lorenzoni, 1971: 139
Negrea and Negrea, 1971: 88, 89, 119
Verner, 1971: n/a, figs 3, 16
Legg, 1972b: 4, 5
Howes, 1972a: 108 (as Chthonius ischonocheles [sic])
Salmon, 1972: 66, 67
Legg, 1973a: 368–392, figs 1–11, plates 1–3
Legg, 1973b: 430–440, figs 1, 2a, plates 1–3
Jones, 1975a: 87–88
Klausen, 1975: 63 (as Chthonius iscnocheles [sic])
Legg, 1975a: 66
Legg, 1975b: 100, figs 1a, 2, 7a, 9a–c, 10a–b, 11a–b
Wood, 1975: 145–149, figs 1–2
Beier, 1977b: 2
Savory, 1977: n/a, fig. 93b
Bellés, 1978: 87
Jones, 1978: 93, 94, 95
Paoletti, 1978: 90
Wood and Gabbutt, 1978: 176–182, figs 6–10, 21–25
Baccetti, 1979: n/a, fig. 11.7
Jones, 1979a: 200–201
Judson, 1979a: 60
Matsuda, 1979: 201 (as Chthonius ischnocheleus [sic])
Rundle, 1979: 47
Schawaller, 1979b: 13
Thaler, 1979: 51
Wood and Gabbutt, 1979a: 285–292
Wood and Gabbutt, 1979b: 329–336
Gabbutt and Aitchison, 1980: 57–60, figs 1–4
Jones, 1980c: n/a, map 3
Judson, 1980: 8
Mahnert, 1981c: n/a, fig. 1
Mendel, 1981: 227
Pieper, 1981: 2
Schawaller, 1981b: 42–44
Mendel, 1982: 296
Harvey, 1983: 126
Jones, 1983: 396
Bacallado, 1984: 110
Jones, 1985: 69
Andersson, Coulianos, Ehnström, Hammarstedt, Imby, Janzon and Waldén, 1987: 68
Ćurčić and Dimitrijević, 1987: 1–2, figs 1a–b
Kovoor, 1987: 161, 166
Ćurčić and Dimitrijević, 1987: 1–2, figs 1a–b
Schaefer, 1988: 100
Callaini and Dallai, 1989: 87, figs 1–3
Chater, 1989: 7
Weygoldt, 1990: 99
Ćurčić and Dimitrijević, 1991b: 91–94
Geoffroy, 1991: 36–37, fig. 2
Pieper, 1991: 218
Gärdenfors and Wilander, 1992: 28, fig. 3a
Harvey, 1992: 150
Harvey, 1992c: n/a, figs 14–17
Peters, 1992: n/a, fig. 810 (as Chthonius ishnocheles [sic])
Callaini and Dallai, 1994: 36, figs 1, 2, 5–12
Fowles, 1994: 17
Poinar and Ćurčić, 1994: 95–96
Ćurčić, Dimitrijević, Makarov, Lučić and Ćurčić, 1996: 45
Legg and O'Connor, 1997: 107–109, fig. 2
Dimitrijević, 1998b: 32–34
Legg, 1998: 2
Proctor, 1998: 163 (as Cthonius [sic] ischnocheles)
Bell, Gates, Haughton, Macdonald, Smith, Wheater and Cullen, 1999: 237–239
Bell, Haughton and Boatman, 1999: 6–7, figs 1–2
Ducháč, 1999a: 14
Wilde, 2001: 3
Krumpál and Krumpálova, 2003: 119
Legg, 2003a: 1
Dimitrijević, 2004b: 17–25, 76–84, figs 1a–d, 2a–d, 3a–d, 4a–d, 5a–d, 29, 30a–h, 31, 32a–I, 33a–c, 34a–I, 35a–e, 36a–l, 37a–c
Negri, 2004: 586–588
Smithers, Thomas and Goodacre, 2005: 3
Stol, 2005: 5, 29, figs 4–6, map 1
Gardini, 2007: 384
Kárpáthegyi, 2007b: 91
Holmen and Scharff, 2008: 68
Kunt, Bayram, Yağmur and Danısman, 2008: 72
Gardini, 2009a: 33
Farkas, Kárpáthegyi, Kiss, Novák and Ujvári, 2009: 63, fig. 4
Arabi, Judson, Deharveng, Lourenco, Cruaud and Hassanin, 2012: n/a, fig. 2A
Cawley, 2016: 117
Legg and Farr-Cox, 2016: n/a, fig. 4
Muster and Blick, 2016: 548, 551
Andrews, 2017b: n/a, fig. 2
Judson, 2018b: 148, figs 5A–B
Christophoryová and Krajčovičová, 2019: 195
Ćurčić, Dimitrijević and Ćurčić, 2020: 18
Gardini, 2021: 75–88, figs 237–265, 407
Muster, Spelda, Rulik, Thormann, von der Mark and Astrin, 2021: 13819, fig. 4
Henrard, Baert, de Smedt, Gardini, Vanherckes, Jocqué, Oger, Kekenbosch, van Nieuwenhove, Lock and Drumont, 2022: 132, 137

Chthonius orthodactylus (Leach):

Morley, 1930: 140 (misidentification)
Morley, 1940: 166 (misidentification)

Chthonius (Chthonius) ischnocheles (Hermann):

Beier, 1932a: 48, fig. 52
Beier, 1934b: 283–285
Leruth, 1935: 26
Bristowe, 1938: 212
Beier, 1939f: 161
Vachon, 1940g: 141
Cooreman, 1946b: 2
Leleup, 1948: 8
Beier, 1949: 2, fig. 1 (as Chtonius [sic] (Chthonius) ischnocheles)
Hoff, 1949b: 433, fig. 13
Hoff, 1951: 2
Beier, 1952e: 2
Beier, 1953a: 293
Evans and Browning, 1954: 8, figs 3, 6–7
Beier, 1955n: 88
Beier, 1958b: 27
Beier, 1958c: 137
Hoff, 1958: 3
Beier, 1959f: 113
Beier, 1961a: 67–68
Beier, 1961b: 21
Beier, 1963a: 253–254
Beier, 1963b: 25, fig. 16
Beier, 1963h: 149
Beier, 1963i: 283
Beier, 1965c: 375
Beier, 1966h: 175
Capolongo, 1969: 202
Hammen, 1969b: 16–17
Beier, 1970a: 45
Legg, 1971a: 475, fig. 1b
Lagar, 1972b: 46
Beier, 1976a: 23
Crocker, 1976: 7–8
Mahnert, 1977c: 62
Crocker, 1978: 8
Caruso and Costa, 1979: 433
Leclerc and Heurtault, 1979: 239
Krumpál, 1980: 23
Callaini, 1982a: 17, 18, fig. 1a
Gardini, 1981a: 102
Krumpál and Kiefer, 1981: 128
Leclerc, 1981: 127–128, plate 2
Anonymous, 1983a: 27
Hammen, 1983a: 10
Heurtault, 1986c: 20
Mahnert, 1986d: 74, fig. 1
Legg and Jones, 1988: 65–66, figs 1–2, 3a–c, 4a–b, 5c, 12a, 12b(a–e)
Capolongo, 1989: 824, 825
Mahnert, 1989a: 85
Mahnert, 1989b: 41
Judson, 1990a: n/a, fig. 1
Gardini, 1991a: 168–169
Harvey, 1991a: 149–151
Gardini and Oggianu, 1992: 13–21, figs 1–11
Hansen, 1992: 105
Mahnert, 1993b: 989
Caoduro, Osella and Ruffo, 1994: 34
Gardini, 1995a: 2
Platen, Blick, Bliss, Drogla, Malten, Martens, Sacher and Wunderlich, 1995: 54
Ashmole, Oromí, Ashmole and Martín, 1996: 18
Gardini, 1996: 159, 161
Petrov, 1997: 262
Austin, Gibson and Harvey, 1998: 346
García Carrillo, 1998: 11
Henderickx, 1999a: 65
Henderickx, 1999b: 67, 68
Gardini, 2000: 17–21
Delfosse, 2003b: 27–28
Gardini, 2004: 125
Ćurčić, Dimitrijević and Legakis, 2004: 233–234
Drogla, 2004: 6
Drogla and Lippold, 2004: 14–15, fig. 2.1
Iorio, 2004: 13
Mahnert, 2004: 463
Ozimec, 2004: 385
Tooren, 2005b: 91–92
Vadell and Zaragoza, 2005: 190–191, fig. 1
Vadell, Zaragoza, Jordana, García, Gràcia and Clamor, 2006: 80–82, fig. 7
Kárpáthegyi, 2007a: 82
Zaragoza, 2007a: 12–13
Zaragoza, Mas and Ribera, 2007: 74–75, figs 1, 3, 5, 7, 9, 11, map 3
Gardini, 2008a: 630
Gardini, 2008b: 4–5
Buddle, 2010: 5
Turienzo, Iorio and Mahnert, 2010: 560, 570, 586
Christophoryová, Štáhlavský and Fedor, 2011: 36, fig. 1M
Christophoryová, Štáhlavský, Krumpál and Fedor, 2012: 4–5
Mahnert and Weber, 2013: 109, figs 1, 2
Zaragoza and Vadell, 2013: 38–39, fig. 3
Kotrbová, Opatova, Gardini and Šťáhlavský, 2016: 327, 330–332, figs 1C, D
Legg, 2019: 190

Chthonius (Chthonius) dacnodes Navás:

Evans and Browning, 1954: 10, figs 10–11 (misidentification)
Legg, 1971a: 475, fig. 1e (misidentification)
Lazzeroni, 1969a: 323–324, fig. 1 (misidentification)

Chthonius (Chthonius) litoralis:

Lazzeroni, 1969b: 393, figs 1–2 (misidentification)

Chthonius (Chthonius) cfr. dacnodes Navás:

Callaini, 1979c: 342–343, figs 2a–f (misidentification)

Not Obisium ischnocheles Hermann:

Théis, 1832: 63–66 (misidentification; see Chthonius orthodactylus (Leach))

Not Chthonius ischnocheles ischnocheles (Hermann):

Callaini, 1986d: 523–524, figs 1a–d (misidentification; see Chthonius rhodochelatus Hadži)


Synonyms
Chthonius rayi dalmatinus Hadži, 1930 -- synonymised by Gardini, 2021: 75
LSID urn:lsid:wac.nmbe.ch:name:cee8c289-ace5-461f-8b50-9b650a306c63
Chthonius rayi dalmatinus
Hadži, 1930a: 72–79, figs 10a–e, 11a–e, 12a–b, 13a–d, 14, 15a–b, 16a–c original description
Hadži, 1930c: 22–24
Chthonius (Chthonius) ischnocheles dalmatinus Hadži:
Beier, 1932a: 48
Chthonius (Chthonius) dalmatinus Hadži:
Hadži, 1933b: 126–130, figs 1a–f
Hadži, 1933c: 173–176
Beier, 1963b: 24
Ćurčić, 1974a: 11
Ćurčić, 1988b: 20–22, figs 14–17
Harvey, 1991a: 146
Ozimec, 2004: 385
Chthonius litoralis Hadži, 1933 -- synonymised by Gardini, 2021: 75
LSID urn:lsid:wac.nmbe.ch:name:e8e60f68-ecad-4dee-843d-ebea7fc90468
Chthonius (Chthonius) litoralis
Hadži, 1933b: 130–133, figs 2a–c original description
Hadži, 1933c: 176–177
Beier, 1963b: 26
Lazzeroni, 1969b: 393, figs 1–2
Ćurčić, 1974a: 12
Ćurčić, 1988b: 19–20, figs 9–13
Harvey, 1991a: 154
Gardini, 2000: 25
Ozimec, 2004: 385
Šťáhlavský and Král, 2004: 51, figs 1b, 5b
Chthonius litoralis Hadži:
Roewer, 1937: 239
Not Chthonius (Chthonius) litoralis:
Lazzeroni, 1969b: 393, figs 1–2 (misidentification; see Chthonius ischnocheles (Hermann))
Chelifer trombidioides Latreille, 1804 -- synonymised by Koch, 1843: 77
LSID urn:lsid:wac.nmbe.ch:name:85d00f19-b6ba-48a6-a7a8-f7edfdab033a
Chelifer trombidioides
Latreille, 1804: 142–143 (synonymised by C.L. Koch, 1843: 77) original description
Latreille, 1806: 133
Leach, 1814: 429 (as Chelifer trombidioidos [sic])
Latreille, 1825c: 133
Brébisson, 1827: 263
Guérin-Méneville, 1828: 598–599
Obisium trombidioides (Latreille):
Leach, 1815: 391
Leach, 1816: 433, plate XXIII
Dugès and Edwards, 1836: n/a, plate 20 bis figs 5, 5a–b
n/a, fig. 547 Shipley, 1909 (reference missing)
Not Chthonius trombidioides (Latreille):
Koch, 1843: 76–79, figs 806–807 (misidentification; see Ephippiochthonius tetrachelatus)
Koch, 1873: 49–50 (misidentification; see Ephippiochthonius tetrachelatus)
Koch, 1877: 180 (misidentification; see Ephippiochthonius tetrachelatus)
Stecker, 1874a: 238 (misidentification; see Ephippiochthonius tetrachelatus)
Stecker, 1875d: 88 (misidentification; see Ephippiochthonius tetrachelatus)
Daday, 1880a: 193 (misidentification; see Ephippiochthonius tetrachelatus)
Tömösváry, 1883: 238–239, plate 5 figs 9–12 (misidentification; see Ephippiochthonius tetrachelatus)
Krauss, 1896: 628 (misidentification; see Ephippiochthonius tetrachelatus)
Chthonius pennsylvanicus Hagen, 1868 -- synonymised by Chamberlin, 1929a: 72
LSID urn:lsid:wac.nmbe.ch:name:a3e591e9-3594-460b-85e3-f60fceddfefd
Chthonius pennsylvanicus
Hagen, 1868b: 52 (as Chthonius pensylvanicus [sic]) (synonymised by Chamberlin, 1929a: 72) original description
Hagen, 1870: 268
Banks, 1890: 152
Banks, 1891: 164
Banks, 1895a: 13
Banks, 1904a: 141
Coolidge, 1908: 114
Ewing, 1911: 80
Pratt, 1927: 410
Not Chthonius pennsylvanicus Hagen:
Berger, 1905: 407 (misidentification; see Tridenchthonius cubanus (Chamberlin))
Chthonius rayi L. Koch, 1873 -- synonymised by Chamberlin, 1929a: 72
LSID urn:lsid:wac.nmbe.ch:name:badbebec-e96e-4a63-a8b9-0b3c18f2f118
Chthonius rayi
Koch, 1873: 48–49 (synonymised by Chamberlin, 1929a: 72) original description
Ritsema, 1874: xxxiv
Stecker, 1874a: 238
Canestrini, 1875a: 2
Stecker, 1875d: 88
Canestrini, 1876: 228
Dale, 1878: 325
Simon, 1879a: 74–75, plate 17 figs 2, 8, plate 19 fig. 19
Cantoni, 1882: 193
Tömösváry, 1883: 240–241, plate 5 figs 13–14
Simon, 1883: 279
Becker, 1884: cclxiv
Canestrini, 1885: n/a, no. 5, figs 1–3
Pickard-Cambridge, 1886: 45
Pickard-Cambridge, 1892: 209–210, plate A figs 10–14
Hansen, 1894: n/a, plate 5 figs 12–13
Carpenter, 1895: 133
Lameere, 1895: 474
Müller and Schenkel, 1895: 822
Becker, 1896: 335–336
Krauss, 1896: 628
Carpenter and Evans, 1897: 123
Ellingsen, 1897: 18–20
Kulczyński, 1899: 458
Tullgren, 1899a: 181
Simon, 1900b: 595
Strand, 1900b: 5
Evans, 1901a: 54
Evans, 1901b: 242
Godfrey, 1901b: 214–215
Goulding, 1901: 154
Kew, 1901b: 194
Donisthorpe, 1902: 67
Woodruffe-Peacock, 1902a: 137
Woodruffe-Peacock, 1902b: 380
Kew, 1903: 298, 299, fig. 2
Planet, 1905: 288
With, 1906: n/a, plate 1 fig. 2a
Ellingsen, 1907a: 172
Godfrey, 1907: 163
Jackson, 1907: 6 (as Chthonius rayii)
Tullgren, 1907e: 246
Carpenter, 1908: 190
Ellingsen, 1908d: 70
Godfrey, 1908: 160
Gozo, 1908: 132
Jackson, 1908: 73 (as Chthonius rayii)
Bignotti, 1909: 67 (as Chthonius ray [sic])
Ellingsen, 1909a: 211
Jackson, 1909: 438 (as Chthonius rayii)
Kew, 1909a: 250
Kew, 1909b: 259
Rossi, 1909: 194
Warburton, 1909: 438
Booth, 1910: 35 (as Chthonius rayii)
Donisthorpe, 1910: 84
Ellingsen, 1910a: 401
Ellingsen, 1910b: 62
Godfrey, 1910: 31–33
Jackson, 1910: 50 (as Chthonius rayii)
Kew, 1910a: 64, 73
Ellingsen, 1911c: 174
Kew, 1911a: 56–57, fig. 19
Lessert, 1911: 37–38, fig. 29
Ellingsen, 1912c: 174
Krausse-Heldrungen, 1912: 65
Oldham, 1912: 299
Standen, 1912: 16, fig. 2
Ellingsen, 1913a: 455
Falcoz, 1913: 3 (as Chtonius [sic] rayi)
Brian, 1914: 11
Kew, 1914b: 93
Kew, 1915: 167
Falconer, 1916: 157–158
Kew, 1916a: 124
Kew, 1916b: 81
Standen, 1916a: 17
Standen, 1916b: 124
Nonidez, 1917: 43
Standen, 1917: 30
Standen, 1918: 335
Navás, 1919: 214
Navás, 1921: 166
Bristowe, 1922: 11
Navás, 1923: 34
Caporiacco, 1925a: 231
Caporiacco, 1926: 75
Navás, 1925: 126, fig. 18b
Jeannel, 1926: 155
Donisthorpe, 1927: 183
Bacelar, 1928: 190
Beier, 1928: 313
Kästner, 1928: 9, fig. 31
Schenkel, 1928: 70, fig. 31
Beier, 1929d: 156
Berland, 1929: 17
Bristowe, 1929a: 25
Bristowe, 1929c: 188
Bristowe, 1929d: 244 (as Chthonius rayii)
Bristowe, 1930: 622 (as Chthonius rayii)
Buresch, 1930: 19
Kew, 1930: 253–256, figs 1b, 2
Bristowe, 1931e: 113 (as Chthonius rayii)
Franganillo Balboa, 1936: 155 (misidentification?)
Morley, 1940: 167
Sanfilipo, Timossi and Conci, 1943: 312 (as Chthonius reyi [sic])
Walton, 1944: 131 (as Chthonius rayii)
Gilbert, 1949: 17
Weidner, 1959: 115
Judson, 1997a: 36–37 (as junior synonym of Chthonius (Chthonius) ischnocheles (Hermann))
Chthonius (Chthonius) rayi L. Koch:
Ellingsen, 1905d: 12
Not Chthonius rayi L. Koch:
Simon, 1898d: 22–23 (misidentification; see Chthonius tenuis L. Koch)
Chthonius (Chthonius) parvioculatus Beier, 1930 -- synonymised by Gardini and Oggianu, 1992: 14
LSID urn:lsid:wac.nmbe.ch:name:79ef81a7-7dc9-483b-8831-108a10e23d96
Chthonius (Chthonius) parvioculatus
Beier, 1930b: 71 (synonymised by Gardini and Oggianu, 1992: 14) original description
Beier, 1932a: 49–50, fig. 54
Roewer, 1937: 239
Beier, 1942: 131
Sanfilipo, Timossi and Conci, 1943: 312
Franciscolo, 1950: 115–116
Beier, 1953e: 105
Beier, 1963b: 26, fig. 17
Gardini, 1980c: 101, 120, 122, 123
Bologna and Taglianti, 1985: 225, fig. 8
Harvey, 1991a: 158
Chthonius parvioculatus Beier:
Wolf, 1938: 608
Sanfilipo, Timossi and Conci, 1943: 312 (not seen)
Franciscolo, 1955: 125
Baggini, 1961: 24
Chthonius (Chthonius) tamaninii Caporiacco, 1952 -- synonymised by Gardini and Oggianu, 1992: 14
LSID urn:lsid:wac.nmbe.ch:name:691af0b0-cf58-490e-b2ff-2c49f35aab64
Chthonius (Chthonius) tamaninii
Caporiacco, 1952b: 63–64, fig. 1 original description
Parenzan, 1955: 303
Beier, 1963b: 26–27
Lazzeroni, 1969a: 322
Lazzeroni, 1970a: 207
Gardini, 1980c: 101, 130
Harvey, 1991a: 161 (synonymised by Gardini and Oggianu, 1992: 14)
Chthonius tamaninii Caporiacco:
Baggini, 1961: 24
Obisium megachelum Amary, 1840 -- synonymised by Gardini, 2021:
LSID urn:lsid:wac.nmbe.ch:name:c98e4b54-fad7-4a5f-9329-58183f76ca91
Obisium megachelum
Amary, 1840: 78, plate IV, figs 1b–d original description
Chthonius rhodochelatus Hadži, 1933 -- synonymised by Gardini, 2021: 75
LSID urn:lsid:wac.nmbe.ch:name:a1180f48-c56d-4be9-9273-c2da15354aba
Chthonius (Chthonius) rhodochelatus
Hadži, 1933b: 133–139, figs 3a–c, 4a–b, 5a–c, 6a–e, 7 original description
Hadži, 1933c: 177–179
Ćurčić, 1974a: 13
Gardini, 2004: 126–127, figs 1–7
Ozimec, 2004: 385
Gardini, 2009a: 39–40
Turienzo, Iorio and Mahnert, 2010: 560, 575
Kotrbová, Opatova, Gardini and Šťáhlavský, 2016: 328, 334, fig. 1G
Chthonius rhodochelatus Hadži:
Roewer, 1937: 239
Judson, 2018b: 148, figs 9A–C
Červená, Gardini, Jablonski and Christophoryová, 2021: 8, figs 2A, 3B
Chthonius ischnocheles ischnocheles (Hermann):
Callaini, 1986d: 523–524, figs 1a–d (misidentification)
Chthonius cfr. densedentatus Beier:
Callaini, 1986e: 234–236, figs 1g-n (misidentification)
Chthonius (Chthonius) ischnocheles reductus Beier:
Inzaghi, 1987: 180–181, figs 18–20 (possible misidentification)
Chthonius (Chthonius) cfr. ruffoi Caporiacco:
Gardini, Sabella and Saccone, 1997: 220–221 (misidentification)
Distribution
Distribution table
Andorra
Austria
Belgium
Bulgaria
Croatia
Czechia
Denmark
France (mainland, Corse)
Germany
Greece (mainland)
Hungary
Ireland
Italy (mainland, Sardinia, Sicily)
Luxembourg
Malta
Netherlands
Norway
Poland
Portugal (mainland, Azores, Madeira)
Romania
Serbia
Slovakia
Spain (mainland, Islas Baleares, Islas Canarias)
St. Helena
Sweden
Switzerland
Turkey
UK
US (Connecticut, Florida, Illinois, Minnesota, New Jersey, New York, North Carolina, Pennsylvania)
Type locality: not stated, but presumably near Strasbourg, Bas-Rhin, Alsace, France.

Coordinates

Locality Lat Lon
1 Strasbourg 48.583333 7.75
Type repository
  • undefined Syntypes: 2 adults (BMNH 1913.9.1.807–808); 1 adult (BMNH 1913.9.1.809); 155 adults (BMNH 1913.9.1.810–959); 8 adults (BMNH 1913.9.1.790–796); 34 specimens (BMNH 1913.9.1.960–989)
Zoobank LSID

urn:lsid:zoobank.org:act:2C723074-5710-44AD-93F6-20D780A164D5

References

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Beier, M. (1958c). Eine neue Neobisium-Art aus den Picentinischen Bergen in Süd-Italien. Memorie del Museo Civico di Storia Naturale di Verona 6: 135–137. -- Show included taxa

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Beier, M. (1963h). Pseudoscorpione aus dem Museum "Enrico Caffi" in Bergamo. Rendiconti dell'Instituto Lombardo di Scienze e Lettere 97B: 147–156. -- Show included taxa

Beier, M. (1963i). Appenninische Pseudoscorpione. Memorie del Museo Civico di Storia Naturale di Verona 10: 283–286. -- Show included taxa

Beier, M. (1965c). Über Pseudoskorpione von den Kanaren. Annalen des Naturhistorischen Museums in Wien 68: 375–381. -- Show included taxa

Beier, M. (1966h). Neues über Höhlen-Pseudoscorpione aus Veneto. Atti della Società Italiana di Scienze Naturali, e del Museo Civile di Storia Naturale, Milano 105: 175–178. -- Show included taxa

Beier, M. (1970a). Ergänzungen zur Pseudoskorpionidenfauna der Kanaren. Annalen des Naturhistorischen Museums in Wien 74: 45–49. -- Show included taxa

Beier, M. (1976a). Die Pseudoscorpione der macaronesischen Inseln. Vieraea, Tenerife 5: 23–32. -- Show included taxa

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Bell, J.R., Gates, S., Haughton, A.J., Macdonald, D.W., Smith, H., Wheater, C.P. & Cullen, W.R. (1999). Pseudoscorpions in field margins: effects of margin age, management and boundary habitats. Journal of Arachnology 27: 236–240. -- Show included taxa

Bell, J.R., Haughton, A.J. & Boatman, N.D. (1999). Population structure of the pseudoscorpion Chthonius ischnocheles (Hermann, 1804) in field margins and the effects of the herbicide glyphosphate. Newsletter of the British Arachnological Society 86: 6–7. -- Show included taxa

Bellés, X. (1978). Notas ecológicas sobre la Cova de la Torre (Sant Feliu de Pallarols, Gironá). Incidencia de los niveles de materia organica sobre la diversidad de la faune terrestre.. Speleon 24: 77–91. -- Show included taxa

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Betts, M.M. (1955). The food of titmice in oak woodland. Journal of Animal Ecology 24: 282–323. -- Show included taxa

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Bristowe, W.S. (1931e). The spiders of the island of Grassholm, and some additions to the Skomer Island list (South Wales). Proceedings of the Zoological Society of London 1931: 111–113. -- Show included taxa

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Buresch, I. (1930). [Monthly report of the Bulgarian Entomological Society]. Mitteilungen der Bulgarischen Entomologischen Gesellschaft in Sofia 5: 19–20. -- Show included taxa

Callaini, G. (1979c). Notulae Chernetologicae. III. Gli pseudoscorpioni della Farma (Arachnida).. Redia 62: 339–354. -- Show included taxa

Callaini, G. (1982a). Étude comparative de la membrane pleurale des pseudoscorpions au microscope électronique à balayage. Memorie, Atti della Società Toscana di Scienze Naturali residente in Pisa B 88 supplemento: 16–26. -- Show included taxa

Callaini, G. (1986d). Note sugli pseudoscorpioni raccolti in alcune grotte della Toscana settentrionale (Arachnida) (Notulae Chernetologicae XXV). Redia 69: 523–542. -- Show included taxa

Callaini, G. (1986e). Pseudoscorpioni dell'Italia settentrionale nel Museo Civico di Storia Naturale di Verona (Arachnida). Notulae Chernetologicae XIX. Bollettino del Museo Civico di Storia Naturale, Verona 12: 229–255. -- Show included taxa

Callaini, G. & Dallai, R. (1989). Les spermatozoïdes des Pseudoscorpions: étude comparative et considérations phylogénétiques. Revue Arachnologique 8: 85–97. -- Show included taxa

Callaini, G. & Dallai, R. (1994). The spermatozoon of pseudoscorpions (Arachnida). Bollettino dell'Accademia Gioenia di Scienze Naturali 26: 35–52. -- Show included taxa

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Caoduro, G., Osella, G. & Ruffo, S. (1994). La fauna cavernicola della regione Veronese. Memorie del Museo Civico di Storia Naturale di Verona (IIa serie), Sezione Scienze della Vita (A: Biologica) 11: 1–144. -- Show included taxa

Capolongo, D. (1969). Studio ecologico delle cantine del Napoletano (primo contributo). Bollettino della Societa Entomologica Italiana 99/101: 193–205. -- Show included taxa

Caporiacco, L. di (1925a). Aracnidi della Provincia de Forlí. Memorie della Società Entomologica Italiana 4: 229–258. -- Show included taxa

Caporiacco, L. di (1926). Secondo saggio sulla fauna aracnologica della Carnia e regioni limitrofe. Memorie della Società Entomologica Italiana 5: 70–130. -- Show included taxa

Caporiacco, L. di (1940a). Arachniden aus der Provinz Verona (Norditalien). Folia Zoologica et Hydrobiologica 10: 1–37. -- Show included taxa

Caporiacco, L. di (1949c). Una piccola raccolta aracnologica dei monti di Calabria. Atti del Museo Civico di Storia Naturel di Trieste 17: 132–136. -- Show included taxa

Caporiacco, L. di (1952b). Aracnidi della Grotta di Agnano (Ostuni, Puglie). Bollettino dei Musei e degli Istituti Biologici dell'Università di Genova 24: 63–65. -- Show included taxa

Carpenter, G.H. (1895). Proceedings of Irish societies. Irish Naturalist 4: 133. -- Show included taxa

Carpenter, G.H. (1908). Phalangida and Chernetida. In: Cole, G.A.J. and Praeger, R.L. (ed.) Handbook to the city of Dublin and the surrounding district Ponsonby and Gibbs, Dublin: 189–190. -- Show included taxa

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Caruso, D. & Costa, G. (1979). Ricerche faunistiche ed ecologiche sulle grotte di Sicilia. VI. Fauna cavernicola di Sicilia (Catalogo ragionato). Animalia, Catania 5: 423–513. -- Show included taxa

Chamberlin, J.C. (1929a). A synoptic classification of the false scorpions or chela-spinners, with a report on a cosmopolitan collection of the same. Part 1. The Heterosphyronida (Chthoniidae) (Arachnida-Chelonethida). Annals and Magazine of Natural History (10) 4: 50–80. -- Show included taxa

Chamberlin, J.C. (1931a). The arachnid order Chelonethida. Stanford University Publications, Biological Sciences 7(1): 1–284. -- Show included taxa

Chamberlin, J.C. (1935b). Chelonethida. In: Pratt, H.S. (ed.) A manual of the common invertebrate animals (exclusive of insects) Blakiston Company, Philadelphia and Toronto: 477–481. -- Show included taxa

Chater, A.O. (1989). False-scorpions in Ceredigion, VC 46. Dyfed Invertebrate Group Newsletter 14: 5–10. -- Show included taxa

Christophoryová, J., Štáhlavský, F. & Fedor, P. (2011). An updated identification key to the pseudoscorpions (Arachnida: Pseudoscorpiones) of the Czech Republic and Slovakia. Zootaxa 2876: 35–48. -- Show included taxa

Christophoryová, J., Štáhlavský, F., Krumpál, M. & Fedor, P. (2012). Pseudoscorpions of the Czech Republic and Slovakia: an annotated and revised checklist (Arachnida: Pseudoscorpiones). North-Western Journal of Zoology 8(1): 1–21. -- Show included taxa

Cloudsley-Thompson, J.L. (1956a). Notes on Arachnida, 25. - An unusual case of phoresy by false-scorpions. Entomologist's Monthly Magazine 92: 71. -- Show included taxa

Cloudsley-Thompson, J.L. (1958b). The behaviour of false-scorpions. Naturalist, London 866: 96. -- Show included taxa

Cloudsley-Thompson, J.L. (1959). Notes on Arachnida, 33. - The fauna of Spaunton Moor, Yorkshire, and of the Galloway Hills.. Entomologist's Monthly Magazine 95: 179. -- Show included taxa

Cloudsley-Thompson, J.L. (1968). Spiders, scorpions, centipedes and mites. Pergamon Press, Oxford. [revised edition] -- Show included taxa

Coolidge, K.R. (1908). A list of the North American Pseudoscorpionida. Psyche, Cambridge 15: 108–114. -- Show included taxa

Cooreman, J. (1946b). Note sur les pseudoscorpions de la faune Belge. Bulletin du Musée Royal d'Histoire Naturelle de Belgique 22(2): 1–8. -- Show included taxa

Crocker, J. (1976). False scorpions. Heritage, Quarterly Bulletin of the Loughborough Naturalists' Club 61: 1–11. -- Show included taxa

Crocker, J. (1978). Introduction to false scorpions. British Arachnological Society, Secretary's News Letter 21: 4–9. -- Show included taxa

Ćurčić, B.P.M. (1974a). Arachnoidea. Pseudoscorpiones. In: Catalogus faunae Jugoslaviae Academie Slovène, Ljubljana 3(4): 1–35. -- Show included taxa

Ćurčić, B.P.M. (1988b). Cave-dwelling pseudoscorpions of the Dinaric karst. Slovenska Akademija Znanosti in Umetnosti, Ljubljana. -- Show included taxa

Ćurčić, B.P.M. & Dimitrijević, R.N. (1987). Two segmental anomolies in Chthonius ischnocheles (Hermann) (Chthoniidae, Pseudoscorpiones). Arhiv Bioloskih Nauka, Beograd 39: 1–2. -- Show included taxa

Ćurčić, B.P.M. & Dimitrijević, R.N. (1991b). Teratological variation in Chthonius ischnocheles (Hermann) and C. aff. tetrachelatus (Preyssler) (Chthoniidae, Pseudoscorpiones). Bulletin de la Société Européenne d'Arachnologie, hors série: 90–96. -- Show included taxa

Ćurčić, B.P.M., Dimitrijević, R.N. & Legakis, A. (2004). The pseudoscorpions of Serbia, Montenegro, and the Republic of Macedonia. Institute of Zoology, Faculty of Biology, University of Belgrade, Belgrade, Athens. -- Show included taxa

Ćurčić, B.P.M., Dimitrijević, R.N., Makarov, S.E., Lučić, L.R. & Ćurčić, S.B. (1996). Further report on nematode-pseudoscorpion assocations. Acta Arachnologica 45: 43–46. -- Show included taxa

Daday, E. (1880a). A magyarországi álskorpiók. Értesítö az Erdélyi Múzeum-egyesület Orvos-Természet-Tudományi Szakosztályából 5: 191–193. -- Show included taxa

Dale, C.W. (1878). History of Glanville's Wootton. Hatchards, London. -- Show included taxa

Delfosse, E. (2003b). Catalogue préliminaire des Pseudoscorpions de France métropolitaine (Arachnida Pseudoscorpiones). Bulletin de Phyllie 17: 24–48. -- Show included taxa

Dimitrijević, R.N. (1998b). Postembryonic development in Chthonius ischnocheles (Hermann) (Chthoniidae, Pseudoscorpiones). Acta Zoologica Bulgarica 50: 31–35. -- Show included taxa

Dimitrijević, R.N. (2004b). Postembryonic development and polymorphism in some pseudoscorpions from the families Chthoniidae and Neobisiidae (Pseudoscorpiones, Arachnida). Monographs, Institute of Zoology, Faculty of Biology, University of Belgrade 10: 1–146. -- Show included taxa

Donisthorpe, H. (1902). Notes on the British myrmecophilous fauna (excluding Coleoptera). Entomologist's Record and Journal of Variation 14: 14–18, 37–40, 67–70. -- Show included taxa

Donisthorpe, H. (1910). On the founding of nests by ants; and a few notes on myrmecophiles. Entomologist's Record and Journal of Variation 22: 82–85. -- Show included taxa

Donisthorpe, H. (1927). The guests of British ants, their habits and life-histories. G. Routledge and Sons, London. -- Show included taxa

Drogla, R. (2004). Kommentierte Checkliste der Pseudoskorpione Thüringens (Arachnida, Pseudoscorpiones). In: Hartmann, M. and Bellstedt, R. (ed.) Check-listen Thüringer Insekten und Spinnentiere Thüringer Entomologenvernad e.V., Jena: 5–8. -- Show included taxa

Drogla, R. & Lippold, K. (2004). Zur Kenntnis der pseudoskorpion-fauna von Ostdeutschland (Arachnida, Pseudoscorpiones). Arachnologische Mitteilungen 27–28: 1–54. -- Show included taxa

Ducháč, V. (1999a). The contemporary research of the pseudoscorpion fauna in the Czech Republic. American Arachnology 59: 14. -- Show included taxa

Dugès, A. & Edwards, M. (1836). Arachnides. In: Cuvier, G. (ed.) Le règne animal distribue d'après son organisation Fortin, Masson, Paris 15–16. [3rd edition] -- Show included taxa

Ellingsen, E. (1897). Norske Pseudoscorpioner. Forhandlinger i Videnskabsselskabet i Kristiania 1896(5): 1–21. -- Show included taxa

Ellingsen, E. (1905d). Pseudoscorpions from Italy and southern France conserved in the R. Museo Zoologico in Torino. Bollettino dei Musei di Zoologia e di Anatomia Comparata della R. Università di Torino 20(503): 1–13. -- Show included taxa

Ellingsen, E. (1907a). Notes on pseudoscorpions, British and foreign. Journal of the Quekett Microscopical Club (2) 10: 155–172. -- Show included taxa

Ellingsen, E. (1908d). Über Pseudoskorpione aus West-Deutschland. Sitzungsberichte herausgegeben vom Naturhistorischen Verein Preussischen Rheinlände und Westfalens 1908: 69–70. -- Show included taxa

Ellingsen, E. (1909a). Contributions to the knowledge of the pseudoscorpions from material belonging to the Museo Civico in Genova. Annali del Museo Civico di Storia Naturale di Genova (3) 4: 205–220. -- Show included taxa

Ellingsen, E. (1910a). Die Pseudoskorpione des Berliner Museums. Mitteilung aus dem Zoologischen Museum in Berlin 4: 357–423. -- Show included taxa

Ellingsen, E. (1910b). Pseudoskorpione und Myriopoden des Naturhistorischen Museums der Stadt Wiesbaden. Jahrbuch des Nassauischen Vereins für Naturkunde 63: 62–65. -- Show included taxa

Ellingsen, E. (1911c). Pseudoscorpionina. In: Le Roi, O. , Zur Fauna des Vereinsgebietes. Sitzungsberichte herausgegeben vom Naturhistorischen Verein Preussischen Rheinlände und Westfalens 1911: 173–174. -- Show included taxa

Ellingsen, E. (1912c). Pseudoscorpiones (troisième série). Archives de Zoologie Expérimentale et Générale (5) 10: 163–175. -- Show included taxa

Ellingsen, E. (1913a). Note on some pseudoscorpions in the British Museum. Annals and Magazine of Natural History (8) 11: 451–455. -- Show included taxa

Evans, G.O. & Browning, E. (1954). Synopses of the British Fauna. No. 10. Pseudoscorpiones. Linnean Society, London. -- Show included taxa

Evans, W. (1901a). Roncus cambridgii, L. K., and other chernetids in Scotland.. Annals of Scottish Natural History 10: 53–54. -- Show included taxa

Evans, W. (1901b). Chthonius tetrachelatus, Preyss., and other chernetids in Scotland. Annals of Scottish Natural History 10: 241. -- Show included taxa

Ewing, H.E. (1911). Notes on pseudoscorpions; a study on the variations of our common species, Chelifer cancroides Linn., with systematic notes on other species. Journal of the New York Entomological Society 19: 65–81. -- Show included taxa

Fage, L. (1933). Les Arachnides cavernicoles de Belgique. Bulletin de la Société Entomologique de France 38: 53–56. -- Show included taxa

Falconer, W. (1916). The harvestmen and pseudoscorpions of Yorkshire. Naturalist, London 41: 103–106, 135–140, 155–158, 191–193. -- Show included taxa

Falcoz, L. (1913). La recherche des arthropodes dans les terriers (fin). Feuille des Jeunes Naturalistes (5) 43: 1–5. -- Show included taxa

Farkas, S., Kárpáthegyi, P., Kiss, M., Novák, J. & Ujvári, Z. (2009). Adatok a Zselic talajlakó mezo- és makrofaunájának ismeretéhez (Nematoda, Pseudoscorpiones, Acari, Chilopoda, Isopoda). Natura Somogyiensis 13: 57–72. -- Show included taxa

Fowles, A. (1994). A provisional checklist of the invertebrates recorded from Wales. 1. False scorpions, harvestmen and spiders (Arachnida). Cyngor Cefn Gwlad Cymru (Countryside Council for Wales), Bangor, Gwynedd. -- Show included taxa

Franciscolo, M. (1950). La fauna della Tann-a do Balou no. 11. Li e descrizione della larva di Bathysciola pumilio Reitt.. Annali del Museo Civico di Storia Naturale di Genova 64: 111–129. -- Show included taxa

Franciscolo, M. (1955). Fauna cavernicola del Savonese. Annali del Museo Civico di Storia Naturale di Genova 67: 1–223. -- Show included taxa

Franganillo Balboa, P. (1936). Los Arácnidos de Cuba hasta 1936. Cultural S.A., La Habana. -- Show included taxa

Gabbutt, P.D. (1967). Quantitative sampling of the pseudoscorpion Chthonius ischnocheles from beech litter. Journal of Zoology, London 151: 469–478. -- Show included taxa

Gabbutt, P.D. (1969c). Life-histories of some British pseudoscorpions inhabiting leaf litter. In: Sheals, J.G. (ed.) The soil ecosystem Systematics Association Publication no. 8, London: 229–235. -- Show included taxa

Gabbutt, P.D. (1970c). Sampling problems and the validity of life history analyses of pseudoscorpions. Journal of Natural History 4: 1–15. -- Show included taxa

Gabbutt, P.D. & Aitchison, C.W. (1980). The effect of temperature and season on the number of hibernation chambers built by adult pseudoscorpions. In: Gruber, J. (ed.) Verhandlungen des 8. Internationalen Arachnologen-Kongress, Wien H. Egermann, Wien: 57–60. -- Show included taxa

Gabbutt, P.D. & Vachon, M. (1963). The external morphology and life history of the pseudoscorpion Chthonius ischnocheles (Hermann). Proceedings of the Zoological Society of London 140: 75–98. -- Show included taxa

García Carrillo, J. (1998). Los Pseudoscorpiones de Aragón (Arachnida: Pseudoscorpionida). Catalogus de la Entomofauna Aragonesa 18: 11–17. -- Show included taxa

Gärdenfors, U. & Wilander, P. (1992). Sveriges klokrypare med nyckel till arterna. Entomologisk Tidskrift 113: 20–35. -- Show included taxa

Gardini, G. (1980c). Catalogo degli Pseudoscorpioni cavernicoli italiani (Pseudoscorpioni d'Italia VIII). Memorie della Società Entomologica Italiana 58: 95–140. -- Show included taxa

Gardini, G. (1981a). Pseudoscorpioni cavernicole Sardi. I. Chthoniidae (Pseudoscorpioni d'Italia, X). Revue Arachnologique 3: 101–114. -- Show included taxa

Gardini, G. (1991a). Pseudoscorpioni cavernicoli del Veneto (Arachnida). (Pseudoscorpioni d'Italia XIX). Bollettino del Museo Civico di Storia Naturale, Verona 15: 167–214. -- Show included taxa

Gardini, G. (1995a). Arachnida Pseudoscorpionida. In: Minelli, A, Ruffo, S. and La Posta, S. (ed.) Checklist delle specie della fauna Italiana Calderini, Bologna 22: 1–8. -- Show included taxa

Gardini, G. (1996). Pseudoscorpioni cavernicoli Italiani seconda nota (Pseudoscorpioni d'Italia XXXI). Bollettino del Museo Civico di Storia Naturale, Verona 20: 157–174. -- Show included taxa

Gardini, G. (2000). Catalogo degli Pseudoscorpioni d'Italia (Arachnida). Fragmenta Entomologica 32, Supplemento: 1–181. -- Show included taxa

Gardini, G. (2004). Note faunistiche e tassonomiche sugli pseudoscorpioni (Arachnida) epigei (Pseudoscorpioni d'Italia XXXV). In: Il Monte Pastello. Memorie del Museo Civico di Storia Naturale di Verona, 2. serie, Monografie Naturalistiche 1: 123–133. -- Show included taxa

Gardini, G. (2007). Order Pseudoscorpiones. In: Gianluca, N. and Vincenzo, V. (ed.) Artopodi del Parco Nazionale del Vesuvio Cierre Edizioni, Sommacampagna: 384–385. -- Show included taxa

Gardini, G. (2008a). Ricerche zoologiche della nave oceanografica “Minerva” (C.N.R.) sulle isole circumsarde. XXXI.. Annali del Museo Civico di Storia Naturale "G. Doria", Genova 99: 629–651. -- Show included taxa

Gardini, G. (2008b). Pseudoscorpioni d’Italia XLII. Chthoniidae cavernicoli di Sardegna (Pseudoscorpiones). Memorie della Società Entomologica Italiana 87: 3–31. -- Show included taxa

Gardini, G. (2009a). Chthonius (C.) delmastroi n. sp. delle Alpi occidentali e del Piemonte e ridescrizione di Chthonius (C.) tenuis L. Koch, 1873 e di C. (C.) submontanus Beier, 1963 (Pseudoscorpiones Chthoniidae). Rivista Piemontese di Storia Naturale 30: 25–51. -- Show included taxa

Gardini, G. & Oggianu, A. (1992). Note tassonomiche e faunistiche su alcuni Chthonius della fauna Italiana (Pseudoscorpionida, Chthoniidae). Fragmenta Entomologica 24: 13–34. -- Show included taxa

Gardini, G., Sabella, G. & Saccone, L. (1997). Studi sulle comunita' di Pseudoscorpioni dei sistemi forestali dei Nebrodi (Sicilia nordorientale) (Pseudoscorpionida). Fragmenta Entomologica 29: 213–237. -- Show included taxa

Geoffroy, J.J. (1991). Les arthropodes cavernicoles de la Grotte du Foulon (Eure-et-Loir, France). In: Aubigny, G.S. (ed.) Actes du 12e Congrès Régional de Spéléologie Région Centre N, Aubigny-sur-Nère 20-21 Octobre 1991 Groupe Spéléologique d'Aubigny-sur-Nère, France, Aubigny-sur-Nère: 35–40. -- Show included taxa

George, R S. (1957). A brief list of the harvestmen (Opiliones) and falsescorpions (Pseudoscorpiones) of Gloucestershire. Proceedings of the Cotteswood Naturalists Field Club 32: 79–81. -- Show included taxa

George, R.S. (1955). Some additional records of Arachnida from Gloucestershire, including four species of mites new to Great Britain. Entomologist's Monthly Magazine 91: 121–124. -- Show included taxa

George, R.S. (1961). More records of Gloucestershire falsescorpions. Report of the North Gloucestershire Naturalists' Society 1959–1960: 38. -- Show included taxa

Gervais, P. (1844). Histoire Naturelle des Insectes. Aptères. Librairie Encyclopédique de Roret, Paris 3. -- Show included taxa

Gestro, R. (1887). Gli Anophthalmus trovati finora in Liguria. Annali del Museo Civico di Storia Naturale di Genova (2) 5: 487–508. -- Show included taxa

Gilbert, O. (1949). Some spiders new to Suffolk. ii. Transactions of the Suffolk Naturalists' Society 7: 17. -- Show included taxa

Gilbert, O. (1951a). Observations on the feeding of some British false scorpions. Proceedings of the Zoological Society of London 121: 547–555. -- Show included taxa

Godfrey, R. (1901b). Chernetidae or false scorpions of West Lothian. Annals of Scottish Natural History 10: 214–217. -- Show included taxa

Godfrey, R. (1907). The false-scorpions of the west of Scotland. Annals of Scottish Natural History 16: 162–163. -- Show included taxa

Godfrey, R. (1908). The false scorpions of Scotland. Annals of Scottish Natural History 17: 90–100, 155–161. -- Show included taxa

Godfrey, R. (1910). The false scorpions of Scotland (continued). Annals of Scottish Natural History 19: 23–33. -- Show included taxa

Gottstein Matočec, S., Bakran-Petricioli, T., Bedek, J., Bukovec, D., Buzjak, S., Franičević, M., Jalžić, B., Kerovec, M., Kletečki, E., Kralj, J., Kružić, P., Kučinić, M., Kuhta, M., Matočec, N., Ozimec, R., Rađa, T., Štamol, V., Ternjej, I. & Tvrtković, N. (2002). An overview of the cave and interstitial biota of Croatia. Natura Croatica 11 (Supplement 1): 1–112. -- Show included taxa

Goulding, R.W. (1901). Lincolnshire Naturalists' Union at Mablethorpe. Naturalist, London 26: 151–154. -- Show included taxa

Gozo, A. (1908). Gli aracnidi di caverne italiane. Bollettino della Societa Entomologica Italiana 38: 109–139. -- Show included taxa

Guérin-Méneville, F.E. (1828). Pince. Chelifer. In: Dictionnaire classique d'histoire naturelle Rey et Gravier, Paris 13: 596–599. -- Show included taxa

Hadži, J. (1930a). Prirodoslovna istrazivanja sjevernodalmatinskog otocja. I. Dugi i kornati. Pseudoscorpiones. Prirodoslovna Istraživanja Kraljevine Jugoslavije 16: 65–79. -- Show included taxa

Hadži, J. (1930c). Pseudoscorpiones. Bulletin International de l'Académie Yugoslave des Sciences et des Beaux-arts 1930: 19–24. -- Show included taxa

Hadži, J. (1933b). Prinos poznavanju pseudoskorpijske faune Primorja. Prirodoslovna Istraživanja Kraljevine Jugoslavije 18: 125–192. -- Show included taxa

Hadži, J. (1933c). Beitrag zur Kenntnis der Pseudoskorpionen-Fauna des Küstenlandes. Bulletin International de l'Académie Yugoslave des Sciences et des Beaux-arts 27: 173–199. -- Show included taxa

Hagen, H. (1868b). The American pseudo-scorpions. Record of American Entomology for the Year 1868: 48–52. -- Show included taxa

Hagen, H. (1870). Synopsis pseudoscorpionidum synonymica. Proceedings of the Boston Society of Natural History 13: 263–272. -- Show included taxa

Hammen, L. van der (1969b). Bijdrage tot de kennis van de Nederlandse bastaardschorpioenen (Arachnida, Pseudoscorpionida). Zoologische Bijdragen 11: 15–24. -- Show included taxa

Hammen, L. van der (1983a). De Spinachtigen (Arachnida) van de ondergrondse kalksteengroeven in Zuid-Limburg. Zoologische Bijdragen 29: 5–51. -- Show included taxa

Hansen, H. (1992). Über die Arachniden-fauna von urbanen lebensräumen in Venedig - II. Die rinde-bewohnenden Arten des stammbereiches von Platanus hybrida (Arachnida: Scorpiones, Pseudoscorpiones, Araneae). Bollettino del Museo Civico di Storia Naturale di Venezia 41: 91–108. -- Show included taxa

Hansen, H.J. (1894). Organs and characters in different orders of arachnids [second part]. Entomologiske Meddelelser 4: 145–249. -- Show included taxa

Harvey, M.S. (1991a). Catalogue of the Pseudoscorpionida. Manchester University Press, Manchester. -- Show included taxa

Harvey, M.S. (1992c). The phylogeny and classification of the Pseudoscorpionida (Chelicerata: Arachnida). Invertebrate Taxonomy 6: 1373–1435. -- Show included taxa

Harvey, P.R. (1983). The arachnids (spiders, harvestmen and pseudoscorpions). In: The Roman River Valley: 122–126. -- Show included taxa

Harvey, P.R. (1992). Pseudoscorpions. Essex Naturalist 11: 149–150. -- Show included taxa

Helversen, O. von (1966a). Pseudoskorpione aus dem Rhein-Main-Gebiet. Senckenbergiana Biologica 47: 131–150. -- Show included taxa

Helversen, O. von (1966b). Über die Homologie der Tasthaare bei Pseudoskorpionen (Arach.). Senckenbergiana Biologica 47: 185–195. -- Show included taxa

Helversen, O. von (1968). Troglochthonius doratodactylus n. sp., ein troglobionter Chthoniide (Arachnida: Pseudoscorpiones: Chthoniidae). Senckenbergiana Biologica 49: 59–65. -- Show included taxa

Henderickx, H. (1999a). Microbisium suecicum: een nieuwe soort voor de Belgische fauna (Arachnida, Pseudoscorpiones). Phegea 27: 1–3. -- Show included taxa

Henderickx, H. (1999b). Naamlijst van de Belgische pseudoschorpioenen (Arachnida Pseudoscorpionida). Bulletin de la Societe Royale Belge d'Entomologie 135: 66–71. -- Show included taxa

Hermann, J.F. (1804). Mémoire aptérologique. F.G. Levrault, Strasbourg. -- Show included taxa

Heurtault, J. (1986c). Pseudoscorpions cavernicoles de France: revue synoptique. Mémoires de Biospéologie 12: 19–32. -- Show included taxa

Hoff, C.C. (1949b). The pseudoscorpions of Illinois. Bulletin of the Illinois Natural History Survey 24: 407–498. -- Show included taxa

Hoff, C.C. (1951). New species and records of chthoniid pseudoscorpions. American Museum Novitates 1483: 1–13. -- Show included taxa

Hoff, C.C. (1958). List of the pseudoscorpions of North America north of Mexico. American Museum Novitates 1875: 1–50. -- Show included taxa

Holmen, M. & Scharff, N. (2008). Anthrenochernes stellae Lohmander, 1939 - status in Denmark for a new species on the EC Habitats Directive (Arachnida, Pseudoscorpiones). Entomologiske Meddelelser- Entomologisk Forening Kobenhavn 76: 55–68. -- Show included taxa

Howes, C.A. (1972a). A review of Yorkshire pseudoscorpions. Naturalist, Hull 918: 107–110. -- Show included taxa

Hunt, S. & Legg, G. (1971). Characterization of the structural protein component in the spermatophore of the pseudoscorpion Chthonius ischnocheles (Hermann). Comparative Biochemistry and Physiology (B) 40: 475–479. -- Show included taxa

Inzaghi, S. (1987). Una nuova specie del genere Chthonius s. str. delle preapli Lombarde. Natura Bresciana 23: 165–182. -- Show included taxa

Iorio, E. (2004). A propos de deux espèces de Pseudoscorpions (Arachnida) présentes en Lorraine. Bulletin de la Société Entomologique de Mulhouse 60: 13–15. -- Show included taxa

Jackson, A.R. (1907). On some rare arachnids captured during 1906. Proceedings of the Chester Society of Natural Science 6: 1–7. -- Show included taxa

Jackson, A.R. (1908). On some rare arachnids captured during 1907. Transactions of the Natural History Society of Northumberland, Durham, and Newcastle-upon-Tyne, new series 3: 49–78. -- Show included taxa

Jackson, A.R. (1909). On some rare arachnids obtained during 1908. Transactions of the Natural History Society of Northumberland, Durham, and Newcastle-upon-Tyne, new series 3: 418–439. -- Show included taxa

Jackson, A.R. (1910). On some arthropods observed in 1909. Lancashire Naturalist (new series) 3: 17–22, 46–51. -- Show included taxa

Jeannel, R. (1926). Faune cavernicole de la France avec une étude des conditions d'existence dans le domaine souterrain. In: Encyclopédique entomologique P. Lechevalier, Paris 7: 1–335. -- Show included taxa

Jones, P.E. (1970a). The occurrence of Chthonius ischnocheles (Hermann) (Chelonethi: Chthoniidae) in two types of hazel coppice leaf litter. Bulletin of the British Arachnological Society 1: 77–80. -- Show included taxa

Jones, P.E. (1975a). The occurrence of pseudoscorpions in the nests of British birds. Proceedings and Transactions of the British Entomological and Natural History Society 8: 87–89. -- Show included taxa

Jones, P.E. (1978). Phoresy and commensalism in British pseudoscorpions. Proceedings and Transactions of the British Entomological and Natural History Society 1978: 90–96. -- Show included taxa

Jones, P.E. (1979a). Pseudoscorpions from Little Wood, Eye, with some additional records from Northamptonshire. Journal of the Northamptonshire Natural History Society and Field Club 37: 198–201. -- Show included taxa

Jones, P.E. (1980c). Provisional atlas of the Arachnida of the British Isles. Part 1. Pseudoscorpiones.. Institute of Terrestrial Ecology, Monks Wood. -- Show included taxa

Jones, R.E. (1983). Some recent Isle of Wight pseudoscorpion records. Proceedings of the Isle of Wight Natural History Archaeological Society 7: 396. -- Show included taxa

Jones, R.E. (1985). The false-scorpions of Norfolk. Transactions of the Norfolk and Norwich Naturalists' Society 27: 67–71. -- Show included taxa

Judson, M.L.I. (1979a). Pseudoscorpions in Hertfordshire. Transactions of the Hertfordshire Natural History Society and Field Club 28: 58–64. -- Show included taxa

Judson, M.L.I. (1980). On some changes in the names of British Chelonethi (Pseudoscorpionida) with a note on the status of Chthonius (C.) dacnodes Navás, 1918, in Britain. Newsletter of the British Arachnological Society 28: 7–9. -- Show included taxa

Judson, M.L.I. (1990a). On the presence of Chthonius (C.) halberti Kew and Chthonius (C.) ressli Beier in France with remarks on the status of Kewochthonius Chamberlin and Neochthonius Chamberlin (Arachnida, Chelonethida, Chthoniidae). Bulletin du Muséum National d'Histoire Naturelle, Paris (4) 11: 593–603. -- Show included taxa

Judson, M.L.I. (1997a). Catalogue of the pseudoscorpion types (Arachnida: Chelonethi) in the Natural History Museum, London. Occasional Papers on Systematic Entomology 11: 1–54. -- Show included taxa

Judson, M.L.I. (2018b). Ontogeny and evolution of the duplex trichobothria of Pseudoscorpiones (Arachnida). Zoologischer Anzeiger 273: 133--151. -- Show included taxa

Kárpáthegyi, P. (2007a). Pseudoscorpions of Hungary. Folia Historico Naturalia Musei Matraensis 31: 81–90. -- Show included taxa

Kárpáthegyi, P. (2007b). Check list of the Hungarian Pseudoscorpiones. Folia Historico Naturalia Musei Matraensis 31: 91–97. -- Show included taxa

Kästner, A. (1928). 2. Ordnung: Moos- oder Afterskorpione, Pseudoscorpiónes Latr. (Chernétes Simon; Chelonéti Thorell; Chernetídea Camb.). In: Brohmer, P, Ehrmann, P. and Ulmer, G. (ed.) Die Tierwelt Mitteleuropas Quelle und Meyer, Leipzig 3 (1, IV): 1–13. -- Show included taxa

Kew, H.W. (1901b). Lincolnshire pseudoscorpions: with an account of the association of such animals with other arthropods. Naturalist, London 26: 193–215. -- Show included taxa

Kew, H.W. (1903). North of England pseudoscorpions. Naturalist, London 28: 293–300. -- Show included taxa

Kew, H.W. (1909a). Notes on the Irish false-scorpions in the National Museum of Ireland. Irish Naturalist 18: 249–250. -- Show included taxa

Kew, H.W. (1909b). Pseudoscorpiones (false-scorpions). In: Grinling, C. et al. (ed.) Survey and record of Woolwich and West Kent South Eastern Union of Scientific Societies, Woolwich: 258–259. -- Show included taxa

Kew, H.W. (1910a). A holiday in south-western Ireland. Notes on some false-scorpions and other animals observed in the counties of Kerry and Cork. Irish Naturalist 19: 64–73. -- Show included taxa

Kew, H.W. (1911a). A synopsis of the false scorpions of Britain and Ireland. Proceedings of the Royal Irish Academy, section B 29: 38–64. -- Show included taxa

Kew, H.W. (1914b). Pseudo-scorpiones. Annual Report and Proceedings of the Belfast Naturalists' Field Club 7: 93. -- Show included taxa

Kew, H.W. (1915). Pseudo-scorpiones. Annual Report and Proceedings of the Belfast Naturalists' Field Club 7: 167. -- Show included taxa

Kew, H.W. (1916a). An historical account of the pseudoscorpion-fauna of the British islands. Journal of the Quekett Microscopical Club (2) 13: 117–136. -- Show included taxa

Kew, H.W. (1916b). A synopsis of the false-scorpions of Britain and Ireland; supplement. Proceedings of the Royal Irish Academy, section B 33: 71–85. -- Show included taxa

Kew, H.W. (1930). On the spermatophores of the Pseudoscorpiones Chthonius and Obisium. Proceedings of the Zoological Society of London 100: 253–256. -- Show included taxa

Klausen, F.E. (1975). Notes on the Pseudoscorpiones of Norway. Norwegian Journal of Entomology 22: 63–65. -- Show included taxa

Koch, C.L. (1843). Die Arachniden. Getreu nach der Natur Abgebildet und Beschrieben. C.H. Zeh'schen, Nürnberg 10. -- Show included taxa

Koch, L. (1873). Uebersichtliche Dartstellung der Europäischen Chernetiden (Pseudoscorpione). Bauer und Raspe, Nürnberg. -- Show included taxa

Koch, L. (1877). Verzeichniss der bei Nürnberg bis jetzt beobachteten Arachniden (mit Ausschluss der Ixodiden und Acariden) und Beschreibungen von neuen, hier vorkommenden Arten. Abhandlungen der Naturhistorischen Gesellschaft zu Nürnberg 6: 113–198. -- Show included taxa

Kotrbová, J., Opatova, V., Gardini, G. & Šťáhlavský, F. (2016). Karyotype diversity of pseudoscorpions of the genus Chthonius (Pseudoscorpiones, Chthoniidae) in the Alps. Comparative Cytogenetics 10: 325–345. -- Show included taxa

Kovoor, J. (1987). Comparative structure and histochemistry of silk-producing organs in arachnids. In: Nentwig, W. (ed.) Ecophysiology of spiders Springer-Verlag, Berlin: 160–186. -- Show included taxa

Krauss, H. (1896). Einiges über Chernetiden nebst einem Auszug der Sammelergebnisse hierüber durch den Entomologischen Verein, Sektion Nürnberg. Illustrierte Wochenschrift für Entomologie 1: 627–628. -- Show included taxa

Krausse-Heldrungen, A.H. (1912). Sardische Chernetiden. Archiv für Naturgeschichte 21: 65–66. -- Show included taxa

Krumpál, M. (1980). Šťúriky (Pseudoscorpionidea) gaderskej doliny (Vel'ká Fatra). Entomologicke Problemy 16: 23–29. -- Show included taxa

Krumpál, M. & Kiefer, M. (1981). Príspevok k Poznaniu Šťúrikov Čel'ade Chthoniidae v ČSSR (Pseudoscorpionidea). Zprávy Československé Společnosti Entomologické Pri CSAV, Praha 17: 127–130. -- Show included taxa

Krumpál, M. & Krumpálova, Z. (2003). Šťúriky - Pseudoscorpiones. In: Mašán, P. and Svatoň, J. (ed.) Pavúkovce Národného Parku Poloniny (Arachnida: Araneae, Pseudoscorpiones, Opiliones, Acari - Parasitiformes) Balada Press, Humenné: 115–126. -- Show included taxa

Kulczyński, V. (1899). Arachnoidea opera Rev. E. Schmitz collecta insulis Maderianis et in insulis Selvages dictis. Rozpravy Akademia Umiejetnosei 36: 319–461. -- Show included taxa

Kunt, K.B., Bayram, A., Yağmur, E.A. & Danısman, T. (2008). Checklist of the pseudoscorpions of Turkey (Pseudoscorpionida; Arachnida). Turkish Journal of Arachnology 1: 70–84. -- Show included taxa

Lagar, A. (1972b). Contribución al conocimiento de los Pseudoescorpiones de España. II. Speleon 19: 45–52. -- Show included taxa

Lameere, A. (1895). Manuel de la faune de Belgique. Lamertin, Bruxelles 1. -- Show included taxa

Latreille, P.A. (1804). Histoire naturelle, generale et particulière, des Crustacés et des Insectes. F. Dufart, Paris 7: 130–132. -- Show included taxa

Latreille, P.A. (1806). Genera crustaceorum et insectorum secundum ordinem naturalem in familias disposita, iconibus exemplisque plurimis explicata. Amand Koenig, Paris 1. -- Show included taxa

Latreille, P.A. (1818b). Tableau encyclopédique et méthodique des trois règnes de la nature. V. Agasse, Paris 24. -- Show included taxa

Latreille, P.A. (1825c). Pince. In: Encyclopédie méthodique. Histoire naturelle. Entomologie, ou histoire naturelle de Crustacés, des Arachnides et des Insectes Agasse, Paris 10: 131–133. -- Show included taxa

Lazzeroni, G. (1969a). Sur la faune de pseudoscorpions de la région apenninique méridionale. (Recherches sur les Pseudoscorpions. III.). Memorie del Museo Civico di Storia Naturale di Verona 16: 321–344. -- Show included taxa

Lazzeroni, G. (1969b). Contributo alla conoscenza degli pseudoscorpioni della regione Veronese. (Ricerche sugli Pseudoscorpioni. IV). Memorie del Museo Civico di Storia Naturale di Verona 16: 379–418. -- Show included taxa

Lazzeroni, G. (1970a). Ricerche sugli Pseudoscorpioni. III. Considerazioni biogeografiche sulla fauna della regione appenninica meridionale. Bulletin du Muséum National d'Histoire Naturelle, Paris (2) 41(supplément 1): 205–208. -- Show included taxa

Leach, W.E. (1814). Crustaceology. In: Brewster, D. (ed.) The Edinburgh Encyclopaedia Blackwood, Edinburgh 7: 383–437. -- Show included taxa

Leach, W.E. (1815). A tabular view of the external characters of four classes of animals, which Linné arranged under Insecta; with the distribution of the genera composing three of these classes into orders, andc. and descriptions of several new genera and species.. Transactions of the Linnean Society of London 11(2): 306–400. doi: https://doi.org/10.1111/j.1096-3642.1813.tb00065.x -- Show included taxa

Leach, W.E. (1816). Annulosa. In: Encyclopedia Britannica Encyclopedia Britannica, London Supplement 1: 401–453. -- Show included taxa

Leclerc, P. (1981). Nouveaux Chthoniidae cavernicoles de la bordure orientale des Cévennes (France) (Arachnides, Pseudoscorpions). Revue Arachnologique 3: 115–131. -- Show included taxa

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Legg, G. (1971a). False-scorpions: the families Chthoniidae, Neobisiidae and Cheiridiidae. Countryside 21: 472–477. -- Show included taxa

Legg, G. (1972b). Some little known arachnids. Newsletter of the British Arachnological Society 4: 4–6. -- Show included taxa

Legg, G. (1973a). Spermatophore formation in the pseudoscorpion Chthonius ischnocheles (Chthoniidae). Journal of Zoology, London 170: 367–394. -- Show included taxa

Legg, G. (1973b). The structure of encysted sperm of some British Pseudoscorpiones (Arachnida). Journal of Zoology, London 170: 429–440. -- Show included taxa

Legg, G. (1975a). A generalised account of the male genitalia and associated glands of pseudoscorpions (Arachnida). Bulletin of the British Arachnological Society 3: 66–74. -- Show included taxa

Legg, G. (1975b). The genitalia and associated glands of five British species belonging to the family Chthoniidae (Pseudoscorpiones: Arachnida). Journal of Zoology, London 177: 99–121. -- Show included taxa

Legg, G. (1998). New species' - what can I find? And others... Try different sampling/trapping techniques. Galea 1: 2. -- Show included taxa

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Legg, G. & Jones, R.E. (1988). Synopses of the British fauna (new series). 40. Pseudoscorpions (Arthropoda; Arachnida). Brill/Backhuys, Leiden. -- Show included taxa

Legg, G. & O'Connor, J.P. (1997). A review of the Irish pseudoscorpions (Arachnida: Pseudoscorpiones). Bulletin of the Irish Biogeographical Society 20: 105–126. -- Show included taxa

Leleup, N. (1947). Contribution a l'étude des Arthropodes nidicoles et microcavernicoles de Belgique. Bulletin et Annales de la Société Entomologique de Belgique 83: 304–343. -- Show included taxa

Leleup, N. (1948). Contribution à l'étude des Arthropodes nidicoles et cavernicoles de Belgique. Mémoires de la Société Entomologique de Belgique 25: 1–55. -- Show included taxa

Leruth, R. (1935). Exploration biologique des cavernes de la Belgique et du Limbourg Hollandais. XXVIIe contribution: Arachnida.. Bulletin du Musée Royal d'Histoire Naturelle de Belgique 11(39): 1–34. -- Show included taxa

Lessert, R. de (1911). Pseudoscorpions. Catalogue des Invertébrés de la Suisse 5: 1–50. -- Show included taxa

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Lucas, H. (1840). Histoire naturelle des Crustacés, des Arachnides et des Myriapodes. P. Duménil, Paris. -- Show included taxa

Mahnert, V. (1977c). Spanische Höhlenpseudoskorpione. Miscelanea Zoologica 4: 61–104. -- Show included taxa

Mahnert, V. (1981c). Sigles trichobothriaux chez les pseudoscorpions (Arachnida). Memorie, Atti della Società Toscana di Scienze Naturali residente in Pisa B 88 supplemento: 185–192. -- Show included taxa

Mahnert, V. (1986d). Arthropodes epigés du Massif de 'San Juan de la Peña' (Jaca, Huesca). Pirineos 124: 73–86. -- Show included taxa

Mahnert, V. (1989a). Les pseudoscorpions (Pseudoscorpiones, Arachnida) récoltés pendant la campagne biospéologique 1987 à Minorque. Endins 14–15: 85–87. -- Show included taxa

Mahnert, V. (1989b). Les pseudoscorpions (Arachnida) des grottes des îles Canaries, avec description de deux espèces nouvelles du genre Paraliochthonius Beier. Mémoires de Biospéologie 16: 41–46. -- Show included taxa

Mahnert, V. (1993b). Pseudoskorpione (Arachnida: Pseudoscorpiones) von Inseln des Mittelmeers und des Atlantiks (Balearen, Kanarische Inseln, Madeira, Ascension), mit vorwiegend subterraner Lebensweise. Revue Suisse de Zoologie 100: 971–992. -- Show included taxa

Mahnert, V. (2004). Die Pseudoskorpione Österreichs (Arachnida, Pseudoscorpiones). Denisia 12: 459–471. -- Show included taxa

Mahnert, V. & Weber, D. (2013). Pseudoskorpione (Arachnida, Pseudoscorpiones) aus Höhlen des Grossherzogtums Luxemburg. Ferrantia 69: 108–114. -- Show included taxa

Marcuzzi, G. (1968). Osservazioni ecologiche sulla fauna del suolo di alcune regioni forestali Italiane. Annali del Centro di Economia Montana delle Venezie 7: 205–331. -- Show included taxa

Marcuzzi, G., Dalla Venezia, L. & Lorenzoni, A.M. (1971). Appunti ecologico-qualitativi sul popolamento animale di alcuni biotopi litorali dell'Alto Adriatico. Atti del Istituto Veneto di Scienze, Lettere ed Arti, Scienze Matematica e Naturali 129: 119–207. -- Show included taxa

Marcuzzi, G., Lorenzoni, A.M. & di Castri, F. (1970). La fauna del suolo di una regione delle Prealpi Venete (M. Spitz, Recoaro), aspetti autoecologici.. Atti del Istituto Veneto di Scienze, Lettere ed Arti, Scienze Matematica e Naturali 128: 411–567. -- Show included taxa

Matsuda, R. (1979). Abnormal metamorphosis and arthropod evolution. In: Gupta, A.P. (ed.) Arthropod phylogeny Van Nostrand Reinhold, New York: 137–256. -- Show included taxa

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Meinertz, T. (1964). Beiträge zur Verbreitung der Pseudoskorpioniden in Dänemark. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i Kjøbenhavn 126: 387–402. -- Show included taxa

Mendel, H. (1981). A review of Suffolk pseudoscorpions. Transactions of the Suffolk Natural History Society 18: 226–232. -- Show included taxa

Mendel, H. (1982). A few more records of pseudoscorpions including a species new to Suffolk. Transactions of the Suffolk Natural History Society 18: 296. -- Show included taxa

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Müller, F. & Schenkel, E. (1895). Verzeichnis der Spinnen von Basel und Umgegend. Verhandlungen der Naturforschenden Gesellschaft in Basel 10: 691–824. -- Show included taxa

Navás, L. (1919). Excursiones entomológicas por Cataluña durante el verano de 1918. Memorias de la Real Academia de Ciencias y Artes de Barcelona (3) 15: 181–214. -- Show included taxa

Navás, L. (1921). Mis excursiones científicas del Verano de 1919. Memorias de la Real Academia de Ciencias y Artes de Barcelona (3) 17: 143–169. -- Show included taxa

Navás, L. (1923). Excursions entomològiques de l'Istiu de 1922. Arxius de l'Institut de Ciències, Barcelona 8: 1–34. -- Show included taxa

Navás, L. (1925). Sinopsis de los Quernetos (Arácnidos) de la Península Ibérica. Broteria, Zoologica 22: 99–130. -- Show included taxa

Negrea, A. & Negrea, Ş. (1971). Sur la synusie du guano des grottes du Banat (Roumanie). Travaux de l'Institute de Spéologie "Émile Racovitza" 10: 81–122. -- Show included taxa

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Parenzan, P. (1953). Fauna del sottosuolo di Napoli (Primo contributo). Bollettino della Società di Naturalisti in Napoli 62: 89–93. -- Show included taxa

Parenzan, P. (1954). Ricerche biologiche nell'Italia meridionale della sezione speleologica dell'I.R.B.. Bollettino della Società di Naturalisti in Napoli 63: 96–101. -- Show included taxa

Parenzan, P. (1955). Speleobiologia dell Murge. Bollettino di Zoologia 22: 293–307. -- Show included taxa

Pearse, A.S. (1943). Chelonethida from the Duke Forest. Anatomical Record 87: 464. -- Show included taxa

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Petrov, B. (1997). A review of Bulgarian pseudoscorpions (Arachnida, Pseudoscorpionida). In: Zabka, M. (ed.) Proceedings of the 16th European Colloquium of Arachnology Wydawnictwo Wyzszej Skoly Rolniczo-Pedagogicznej, Siedlce: 261–269. -- Show included taxa

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Pickard-Cambridge, O. (1892). On the British species of false-scorpions. Proceedings of the Dorset Natural History and Antiquarian Field Club and Archaeological Society 13: 199–231. -- Show included taxa

Pieper, H. (1981). Die Pseudoskorpione von Madeira und nachbarinseln. Bocagiana, Funchal 60: 1–7. -- Show included taxa

Pieper, H. (1991). New records of pseudoscorpions from the Madeiran Islands. Boletim do Museu Municipal do Funchal 42: 217–221. -- Show included taxa

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Platen, R., Blick, T., Bliss, P., Drogla, R., Malten, A., Martens, J., Sacher, P. & Wunderlich, J. (1995). Verzeichnis der Spinnentiere (excl. Acarida) Deutschlands (Arachnida: Araneida, Opilionida, Pseudoscorpionida). Arachnologische Mitteilungen, Sonderband 1: 1–55. -- Show included taxa

Poinar, G.O., Jr & Ćurčić, B.P.M. (1994). Nematode (Nematoda) associates and parasites of pseudoscorpions (Arachnida). Journal of Invertebrate Pathology 63: 95–96. -- Show included taxa

Pontin, A.J. (1961). The prey of Lasius niger (L.) and L. flavus (F.) (Hym., Formicidae). Entomologist's Monthly Magazine 97: 135–137. -- Show included taxa

Pratt, H.S. (1927). A manual of the common invertebrate animals exclusive of insects. A.C. McClurg and Co., Chicago. -- Show included taxa

Proctor, H.C. (1998). Indirect sperm transfer in arthropods: behavioral and evolutionary trends. Annual Review of Entomology 43: 153–174. -- Show included taxa

Ritsema, C. (1874). Naamlijst der tot Heden in Nederland waargenomen bastaard-schorpioenen (Chernetiden). Tijdschrift voor Entomologie 18: xxxiii–xxxiv. -- Show included taxa

Roewer, C.F. (1936). Chelonethi oder Pseudoskorpione. In: Bronns, H.G. (ed.) Bronn's Klassen und Ordnungen des Tierreichs Akademische Verlagsgesellschaft M.B.H., Leipzig 5(IV)(6)(1): 1–160. -- Show included taxa

Roewer, C.F. (1937). Chelonethi oder Pseudoskorpione. In: Bronns, H.G. (ed.) Bronn's Klassen und Ordnungen des Tierreichs Akademische Verlagsgesellschaft M.B.H., Leipzig 5(IV)(6)(1): 161–320. -- Show included taxa

Rossi, A. (1909). Materiali per una fauna aracnologica della Provincia di Roma. Chernetes (Pseudoscorpioni). Bollettino della Societa Entomologica Italiana 40: 182–194. -- Show included taxa

Rundle, A.J. (1979). Pseudoscorpions and harvestmen in Bedfordshire. Bedfordshire Naturalist 33: 47–50. -- Show included taxa

Salmon, S.J. (1972). Recent records of British pseudoscorpions. Bulletin of the British Arachnological Society 2: 66–68. -- Show included taxa

Sanfilipo, N., Timossi, G. & Conci, C. (1943). La grotta del Brigidun e la grotta Dragonara (Esplorazioni speleologiche nella Provincia di Genova - I). Annali del Museo Civico di Storia Naturale di Genova 61: 307–319. -- Show included taxa

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Savory, T.H. (1945). The spiders and allied orders of the British Isles comprising descriptions of every family of British spiders, every species of harvestmen and false scorpions also the more familiar of the British mites and sea-spiders. Frederick Warne & Co., London. [2nd edition] -- Show included taxa

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Schaefer, M. (1988). 1. Chelicerata, Spinnentiere. In: Brohmer, P. (ed.) Fauna von Deutschland Quelle und Meyer, Heidelberg: 99–135. [17th edition] -- Show included taxa

Schawaller, W. (1979b). Einige Pseudoskorpione und Weberknechte aus dem landschaftsschutzgebiet Poppenweiler bei Ludwigsburg. Mitteilungen Entomologischer Verein Stuttgart 14: 12–13. -- Show included taxa

Schawaller, W. (1981b). Pseudoskorpione von Korsika (Arachnida, Pseudoscorpionidea). Entomologica Basiliensia 6: 42–51. -- Show included taxa

Schenkel, E. (1928). Pseudoscorpionida (Afterskorpione). In: Dahl, F. (ed.) Die Tierwelt Deutschlands G. Fischer, Jena 8: 52–72. -- Show included taxa

Schenkel, E. (1938). Die Arthropodenfauna von Madeira nach den Ergebnissen der Reise von Prof. Dr. O. Lundblad Juli-August 1935. IV. Araneae, Opiliones und Pseudoscorpiones. Arkiv för Zoologi 30(7): 1–42. -- Show included taxa

Simon, E. (1879a). Les Ordres des Chernetes, Scorpiones et Opiliones. In: Les Arachnides de France Librairie Encyclopédique de Roret, Paris 7: 1–332. -- Show included taxa

Simon, E. (1883). Matériaux pour servir à la faune arachnologique des îles de l'Océan Atlantique (Açores, Madère, Salvages, Canaries, Cap Vert, Sainte-Hélène et Bermudes). Annales de la Société Entomologique de France (6) 3: 259–314. -- Show included taxa

Simon, E. (1896). Note sur quelques Chernetes de Ligurie. Annali del Museo Civico di Storia Naturale di Genova (2) 16: 372–376. -- Show included taxa

Simon, E. (1898d). Studio sui Chernetes Italiani conservati nel Museo Civico di Genova con descrizione di nuova specie. Annali del Museo Civico di Storia Naturale di Genova (2) 19: 20–24. -- Show included taxa

Simon, E. (1900b). Studio sui Chernetes Italiani conservati nel Museo Civico di Genova. II. Annali del Museo Civico di Storia Naturale di Genova (2) 20: 593–595. -- Show included taxa

Smithers, P., Thomas, G. & Goodacre, S. (2005). The Annual Meeting May 14th-15th 2005. Slapton Downs, Devon. Newsletter of the British Arachnological Society 104: 1–4. -- Show included taxa

Šťáhlavský, F. & Král, J. (2004). Karyotype analysis and achiasmatic meiosis in pseudoscorpions of the family Chthoniidae (Arachnida: Pseudoscorpiones). Hereditas 140: 49–60. -- Show included taxa

Standen, R. (1912). The false-scorpions of Lancashire and some adjoining counties, with a preliminary list of records. Lancashire and Cheshire Naturalist 5: 7–16. -- Show included taxa

Standen, R. (1916a). Report on the false-scorpions and woodlice for 1915–16. Lancashire and Cheshire Naturalist 9: 17–20. -- Show included taxa

Standen, R. (1916b). Chelifer (Chernes) powelli Kew and Chelifer (Withius) subruber Simon in Lancashire and Cheshire. Lancashire and Cheshire Naturalist 9: 124–125. -- Show included taxa

Standen, R. (1917). Report on the false-scorpions (Chelifers). Lancashire and Cheshire Naturalist 10: 27–31. -- Show included taxa

Standen, R. (1918). Report on the false-scorpions (Chelifers) for 1917. Lancashire and Cheshire Naturalist 10: 335. -- Show included taxa

Stecker, A. (1874a). Zur Kenntnis der Chernetidenfauna Böhmens. Sitzungsberichte der Königliche Böhmischen Gesellschaft der Wissenschaften 8: 227–241. -- Show included taxa

Stecker, A. (1875d). Die Chernetidenfauna Böhmens. Zeitschrift für die Gesammten Naturwissenschaften 11: 87–89. -- Show included taxa

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Strand, E. (1900b). Zur Kenntniss der Arachniden Norwegens. Kongelige Norske Videnskabernes Selskabs Skrifter 1900(2): 1–46. -- Show included taxa

Szalay, L. (1968). Pókszabásúak I. Arachnoidea I. In: Magyarország Allatvilága (Faunae Hungariae) Akadémiai Kiadó, Budapest 18(1): 1–122. -- Show included taxa

Thaler, K. (1979). Fragmenta faunistica Tirolensia, IV (Arachnida: Acari: Caeculidae; Pseudoscorpiones; Scorpiones; Opiliones; Aranei; Insecta: Dermaptera; Thysanoptera; Diptera Nematocera: Mycetophilidae, Psychodidae, Limoniidae und Tipulidae). Veröffentlichungen des Museums Ferdinandeum 59: 49–83. -- Show included taxa

Théis, C. de (1832). Lettre adressée à M. Audouin, sur quelques Arachnides des genres Hydrachna et Chelifer. Annales des Sciences Naturelles 27: 57–78. -- Show included taxa

Tömösváry, Ö. (1883). Pseudoscorpiones faunae Hungaricae. A Magyar fauna álskorpiói. Magyar Tudományos Akadémia Matematikai és Természettudományi Közlemények 18: 135–256. -- Show included taxa

Tooren, D. van den (2005b). Naamlijst en determinatiesleutel tot de Pseudoschorpioenen van Nederland (Arachnida: Pseudoscorpiones). Nederlandse Faunistische Mededelingen 23: 91–102. -- Show included taxa

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Tullgren, A. (1907e). Über einige Chelonethiden des Naturhistorischen Museums zu Wiesbaden. Jahrbuch des Nassauischen Vereins für Naturkunde 60: 246–248. -- Show included taxa

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Vachon, M. (1934g). Pseudoscorpionides de la Côte-d'Or (2e liste). Bulletin Scientifique de Bourgogne 4: 162–163. -- Show included taxa

Vachon, M. (1934i). Sur le développement post-embryonnaire des pseudoscorpionides. Deuxiemes note. Les formes immatures de Chelifer cancroides L. (suite). Bulletin de la Société Zoologique de France 59: 405–416. -- Show included taxa

Vachon, M. (1935j). Sur les Pseudoscorpions qui habitent les maisons et leurs dépendances. Bulletin Scientifique de Bourgogne 5: 187–188. -- Show included taxa

Vachon, M. (1936a). Sur le développement postembryonnaire des Pseudoscorpions (quatrième note). Les formules chaetotaxiques des pattes-mâchoires.. Bulletin du Muséum National d'Histoire Naturelle, Paris (2) 8: 77–83. -- Show included taxa

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Vachon, M. (1940g). Éléments de la faune portugaise des Pseudoscorpions (Arachnides) avec description de quatre espèces nouvelles. Anais da Faculdade de Ciencias do Porto Academia Polytechnica do Porto 25: 141–164. -- Show included taxa

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Vachon, M. (1961). Remarques sur les Pseudoscorpions de Madère, des Açores et les Canaries (première note). Bulletin du Muséum National d'Histoire Naturelle, Paris (2) 33: 98–104. -- Show included taxa

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Vadell, M., Zaragoza, J.A., Jordana, R., García, L., Gràcia, F. & Clamor, B. (2006). Nuevas Aportaciones al conocimiento de la fauna cavernícola terrestre de las Coves del Pirata, Cova de Point, Cova de sa piqueta y la cova des Xots (Manacor, Mallorca, Balaeres). Endins 29: 75–98. -- Show included taxa

Verner, P.H. (1971). Štírci - Pseudoscorpionidea. In: Daniel, M. and Černý, V. (ed.) Klič Zviřeny ČSSR Českoslovenká Akademie Věd, Praha 4: 19–31. -- Show included taxa

Walton, G.A. (1944). The natural history of Read's Cave. Proceedings of the University of Bristol Spelaeological Society 5: 127–137. -- Show included taxa

Warburton, C. (1909). Arachnida Embolobranchiata (scorpions, spiders, mites, etc.). In: Harmer, S.F. and Shipley, A.E. (ed.) The Cambridge natural history MacMillan and Co., London 4: 297–474. -- Show included taxa

Weidner, H. (1959). Die Entomologischen Sammlungen des Zoologischen Staatsinstituts und Zoologischen Museums Hamburg. I. Teil. Pararthropoda und Chelicerata I. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institute 57: 89–142. -- Show included taxa

Weygoldt, P. (1966a). Vergleichende Untersuchungen zur Fortpflanzungsbiologie der Pseudoscorpione: Beobachtungen über das Verhalten, die Samenubertragungsweisen und die Spermatophoren einiger einheimischer Arten. Zeitschrift für Morphologie und Ökologie der Tiere 56: 39–92. -- Show included taxa

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Weygoldt, P. (1968). Vergleichend-embryologische Untersuchungen an Pseudoscorpionen. IV. Die Entwicklung von Chthonius tetrachelatus Preyssl., Chthonius ischnocheles (Hermann) (Chthoniinea, Chthoniidae) und Verrucaditha spinosa Banks (Chthoniinea, Tridenchthoniidae). Zeitschrift für Morphologie und Ökologie der Tiere 63: 111–154. -- Show included taxa

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Weygoldt, P. (1990). Arthropoda-Chelicerata: sperm transfer. In: Adiyodi, K.G. and Adiyodi, R.G. (ed.) Reproductive biology of invertebrates Oxford & IBH Publishing Co., New Delhi 4 (B): 77–119. -- Show included taxa

Wilde, I. (2001). Annual general meeting and field weekend Castle Head, Grange-over-Sands, Cumbria 1st-3rd June, 2001. Newsletter of the British Arachnological Society 92: 1–3. -- Show included taxa

With, C.J. (1906). The Danish expedition to Siam 1899–1900. III. Chelonethi. An account of the Indian false-scorpions together with studies on the anatomy and classification of the order. Oversigt over det Konigelige Danske Videnskabernes Selskabs Forhandlinger (7) 3: 1–214. -- Show included taxa

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Wood, P.A. (1975). Cyclical gonadial development of Chthonius ischnocheles (Hermann). In: Proceedings of the 6th International Arachnological Congress Vrije Universiteit of Amsterdam, Amsterdam: 145–149. -- Show included taxa

Wood, P.A. & Gabbutt, P.D. (1978). Seasonal vertical distribution of pseudoscorpions in beech litter. Bulletin of the British Arachnological Society 4: 176–183. -- Show included taxa

Wood, P.A. & Gabbutt, P.D. (1979a). Silken chambers built by adult pseudoscorpions in laboratory culture. Bulletin of the British Arachnological Society 4: 285–293. -- Show included taxa

Wood, P.A. & Gabbutt, P.D. (1979b). Silken chambers built by nymphal pseudoscorpions in laboratory culture. Bulletin of the British Arachnological Society 4: 329–336. -- Show included taxa

Woodruffe-Peacock, E.A. (1902a). Lincolnshire naturalists at Torksey. Naturalist, London 27: 133–138. -- Show included taxa

Woodruffe-Peacock, E.A. (1902b). Lincolnshire naturalists at Scunthorpe. Naturalist, London 27: 375–380. -- Show included taxa

Zangheri, P. (1966). Repertorio sistematico e topografico della flora e fauna vivente e fossile della Romagna. Museo Civico di Storia Naturale di Verona, Memorie Fuori Serie 1 2: 485–854. -- Show included taxa

Zaragoza, J.A. (2007a). Catálogo de los Pseudoescorpiones de la Península Ibérica e Islas Baleares (Arachnida: Pseudoscorpiones). Revista Ibérica de Aracnología 13: 3–91. -- Show included taxa

Zaragoza, J.A. & Vadell, M. (2013). Chthonius (Chthonius) campaneti, a new pseudoscorpion species from Coves de Campanet, Mallorca (Spain). Revista Ibérica de Aracnología 23: 33–41. -- Show included taxa

Zaragoza, J.A., Mas, E. de & Ribera, C. (2007). Pseudoescorpiones del Parque Naturel del Cadí-Moixeró (Pirineo Catalán): estudio ecológico, faunístico y taxonómico (Arachnida: Pseudoscorpiones). Revista Ibérica de Aracnología 14: 69–95. -- Show included taxa

Gardini, G. (2021). The Italian species of the Chthonius ischnocheles group (Arachnida, Pseudoscorpiones, Chthoniidae), with reference to neighbouring countries. Zootaxa 4987: 1‒131. -- Show included taxa

Cawley, M. (2016). An indexed bibliographical checklist of the false-scorpions (Arachnida: Pseudoscorpiones) of Ireland (1836-2014). Bulletin of the Irish Biogeographical Society 40: 116–124. -- Show included taxa

Legg, G. (2019). Changes in the British pseudoscorpion fauna over the last 50 years. Arachnology 18: 189–195. -- Show included taxa

Červená, M., Gardini, G., Jablonski, D. & Christophoryová, J. (2021). Checklist of pseudoscorpions (Arachnida, Pseudoscorpiones) of Albania. Zoological Studies 60(17): 1–25. -- Show included taxa

Christophoryová, J. & Krajčovičová, K. (2019). Faunistic records of pseudoscorpions (Arachnida: Pseudoscorpiones) from South East Poland. Klapalekiana 55: 193–198. -- Show included taxa

Muster, C., Spelda, J., Rulik, B., Thormann, J., von der Mark, L> & Astrin, J.J. (2021). The dark side of pseudoscorpion diversity: The German Barcode of Life campaign reveals high levels of undocumented diversity in European false scorpions. Ecology and Evolution 11(20): 13815–13829. -- Show included taxa

Legg, G. & Farr-Cox, F. (2016). Illustrated key to the British false scorpions (pseudoscorpions). Field Studies Council: Telford. -- Show included taxa

Capolongo, D. (1989). Specie cavernicole di Campania: primo aggiornamento. -- Show included taxa

Ćurčić, N.B., Ćurčić, N.B. & Ćurčić, S.B. (2020). Checklist of the pseudoscorpions (Arachnida: Pseudoscorpiones) of Serbia. Arthropoda Selecta 29: 13–27. -- Show included taxa

Henrard, A., Baert, L., de Smedt, P., Gardini, G., Vanherckes, L., Jocqué, R., Jocqué, R., Kekenbosch, R., van Nieuwenhove, C., Lock, K. & Drumont, A. (2022). On the arachnofauna of the Jean Massart botanical garden (Brussels-Capital Region, Belgium). Journal of the Belgian Arachnological Society 27: 122–137. -- Show included taxa

Muster, C. & Blick, T. (2016). Rote Liste und Gesamtartenliste der Pseudoskorpione (Arachnida: Pseudoscorpiones) Deutschlands.. Naturschutz und Biologische Vielfalt 70: 539–561. -- Show included taxa

Logs
Date Type Reference Old value
2024-02-06 Taxonomic reference update Vachon, 1940
2024-02-01 New taxonomic reference entry Judson, 2018
2023-11-22 New taxonomic reference entry Weygoldt, 1966
2023-04-22 New taxonomic reference entry Muster and Blick, 2016
2023-04-17 New taxonomic reference entry Henrard, Baert, de Smedt, Gardini, Vanherckes, Jocqué, Oger, Kekenbosch, van Nieuwenhove, Lock and Drumont, 2022
2022-12-17 Taxonomic reference update Andrews, 2017
2022-12-17 Taxonomic reference update Andrews, 2017
2022-12-17 New taxonomic reference entry Andrews, 2017
2022-11-18 New taxonomic reference entry Ćurčić, Dimitrijević and Ćurčić, 2020
2022-11-13 New taxonomic reference entry Christophoryová and Krajčovičová, 2019
2022-10-16 New taxonomic reference entry Capolongo, 1989
2022-10-16 New taxonomic reference entry Legg, 2019
2022-10-03 New taxonomic reference entry Legg and Farr-Cox, 2016
2022-09-25 New taxonomic reference entry Muster, Spelda, Rulik, Thormann, von der Mark and Astrin, 2021
2022-06-18 Taxonomic reference update Callaini, 1986
2022-06-18 Taxonomic reference update Théis, 1832
2022-06-18 Taxonomic reference update Pearse, 1943
2022-04-06 New taxonomic reference entry Cawley, 2016
2022-03-20 Species synonymy Hadži, 1933
2022-03-20 Species synonymy Hadži, 1933
2022-03-20 Species synonymy Hadži, 1930
2022-03-20 Species synonymy Amary, 1840
2022-03-20 New taxonomic reference entry Gardini, 2021
2022-03-18 Species updated Old value